1 Physics and Cell Biology Laboratory, Department of Chromosome Science, National Institute of Genetics, Shizuoka, Japan.
2 Department of Genetics, SOKENDAI University, Shizuoka, Japan.
3 Department of Molecular Physiology and Biological Physics, Center for Membrane and Cell Physiology, School of Medicine, University of Virginia, Charlottesville, VA, United States.
4 Department of Cell Biology, School of Medicine, University of Virginia, Charlottesville, VA, United States.
5 Department of Molecular and Cellular Biology and School of Engineering and Applied Sciences, Harvard University, Cambridge, MA, United States.
6 Center for Computational Biology, Flatiron Institute, New York, NY, United States.