1 Oxford University, Weatherall Institute of Molecular Medicine, John Radcliffe Hospital Oxford, Oxford, OX3 9DS, UK.
2 School of Mathematics and Statistics, University of St Andrews, Scotland, KY16 9SS, UK.
3 Institute of Evolutionary Biology, School of Biological Sciences, University of Edinburgh, Charlotte Auerbach Road, Edinburgh, EH9 3FL, UK. [email protected].
4 School of Life Sciences, University of Sussex, Brighton, BN1 9QG, UK.