1 Department of Clinical and Experimental Medicine, Brighton and Sussex Medical School, Brighton, UK.
2 Department of Gastroenterology and Hepatology, Brighton and Sussex University Hospitals NHS Trust, Brighton, UK.
3 NIHR Nottingham Biomedical Research Centre, Nottingham University Hospitals NHS Trust and the University of Nottingham, Nottingham, UK.
4 Nottingham Digestive Diseases Centre, School of Medicine, University of Nottingham, Nottingham, Nottinghamshire, UK.
5 NIHR Nottingham Biomedical Research Centre, Nottingham University Hospitals NHS Trust and the University of Nottingham, Nottingham, UK [email protected].