OX40L/OX40 Signal Promotes IL-9 Production by Mucosal MAIT Cells During Helicobacter pylori Infection

Front Immunol. 2021 Mar 11:12:626017. doi: 10.3389/fimmu.2021.626017. eCollection 2021.

Abstract

Mucosal associated invariant T (MAIT) cells play a critical role in Helicobacter pylori (H. pylori)-induced gastritis by promoting mucosal inflammation and aggravating mucosal injuries (1, 2). However, the underlying mechanism and key molecules involved are still uncertain. Here we identified OX40, a co-stimulatory molecule mainly expressed on T cells, as a critical regulator to promote proliferation and IL-9 production by MAIT cells and facilitate mucosal inflammation in H. pylori-positive gastritis patients. Serum examination revealed an increased level of IL-9 in gastritis patients. Meanwhile, OX40 expression was increased in mucosal MAIT cells, and its ligand OX40L was also up-regulated in mucosal dendritic cells (DCs) of gastritis patients, compared with healthy controls. Further results demonstrated that activation of the OX40/OX40L pathway promoted IL-9 production by MAIT cells, and MAIT cells displayed a highly-activated phenotype after the cross-linking of OX40 and OX40L. Moreover, the level of IL-9 produced by MAIT cells was positively correlated with inflammatory indexes in the gastric mucosa, suggesting the potential role of IL-9-producing MAIT cells in mucosal inflammation. Taken together, we elucidated that OX40/OX40L axis promoted mucosal MAIT cell proliferation and IL-9 production in H. pylori-induced gastritis, which may provide potential targeting strategies for gastritis treatment.

Keywords: H. pylori; IL-9; MAIT cells; OX40; gastritis; inflammation.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Adult
  • Case-Control Studies
  • Cell Proliferation
  • Cells, Cultured
  • Coculture Techniques
  • Female
  • Gastric Mucosa / immunology
  • Gastric Mucosa / metabolism
  • Gastric Mucosa / microbiology*
  • Gastritis / immunology
  • Gastritis / metabolism
  • Gastritis / microbiology*
  • Helicobacter Infections / immunology
  • Helicobacter Infections / metabolism
  • Helicobacter Infections / microbiology*
  • Helicobacter pylori / immunology
  • Helicobacter pylori / pathogenicity*
  • Host-Pathogen Interactions
  • Humans
  • Immunity, Mucosal
  • Interleukin-9 / metabolism*
  • Lymphocyte Activation
  • Male
  • Middle Aged
  • Mucosal-Associated Invariant T Cells / immunology
  • Mucosal-Associated Invariant T Cells / metabolism
  • Mucosal-Associated Invariant T Cells / microbiology*
  • OX40 Ligand / metabolism*
  • Phenotype
  • Receptors, OX40 / metabolism*
  • Signal Transduction
  • Young Adult

Substances

  • IL9 protein, human
  • Interleukin-9
  • OX40 Ligand
  • Receptors, OX40
  • TNFRSF4 protein, human
  • TNFSF4 protein, human