1 Research Centre on Animal Cognition, Centre for Integrative Biology, CNRS, University of Toulouse, 118 Route de Narbonne, 31062 Cedex 09, Toulouse, France.
2 Centre for Integrative Biology, Université de Toulouse, CNRS, University of Toulouse, 118 Route de Narbonne, 31062 Cedex 09, Toulouse, France.
3 Evolution, Genomes, Behavior and Ecology, CNRS (UMR 9191), IRD, University Paris Saclay, 1 Avenue de la Terrasse, Gif-sur-Yvette 91198, France.
4 Latin American Society for Bee Research (SOLATINA), Bolivian Chapter, Santivañez 0134, Cochabamba, Bolivia.