Analysis of Protein Cysteine Acylation Using a Modified Suspension Trap (Acyl-Trap)

J Proteome Res. 2024 Aug 2;23(8):3716-3725. doi: 10.1021/acs.jproteome.4c00225. Epub 2024 Jul 15.

Abstract

Proteins undergo reversible S-acylation via a thioester linkage in vivo. S-palmitoylation, modification by C16:0 fatty acid, is a common S-acylation that mediates critical protein-membrane and protein-protein interactions. The most widely used S-acylation assays, including acyl-biotin exchange and acyl resin-assisted capture, utilize blocking of free Cys thiols, hydroxylamine-dependent cleavage of the thioester and subsequent labeling of nascent thiol. These assays generally require >500 μg of protein input material per sample and numerous reagent removal and washing steps, making them laborious and ill-suited for high throughput and low input applications. To overcome these limitations, we devised "Acyl-Trap", a suspension trap-based assay that utilizes a thiol-reactive quartz to enable buffer exchange and hydroxylamine-mediated S-acyl enrichment. We show that the method is compatible with protein-level detection of S-acylated proteins (e.g., H-Ras) as well as S-acyl site identification and quantification using "on trap" isobaric labeling and LC-MS/MS from as little as 20 μg of protein input. In mouse brain, Acyl-Trap identified 279 reported sites of S-acylation and 1298 previously unreported putative sites. Also described are conditions for long-term hydroxylamine storage, which streamline the assay. More generally, Acyl-Trap serves as a proof-of-concept for PTM-tailored suspension traps suitable for both traditional protein detection and chemoproteomic workflows.

Keywords: Fragpipe; S-Trap; S-acylation; Suspension trapping; Swisspalm; TMT; chemoproteomics S-palmitoylation; cysteine proteomics; isobaric labeling; thioester.

MeSH terms

  • Acylation
  • Animals
  • Brain / metabolism
  • Chromatography, Liquid / methods
  • Cysteine* / chemistry
  • Cysteine* / metabolism
  • Hydroxylamine / chemistry
  • Lipoylation
  • Mice
  • Protein Processing, Post-Translational
  • Proteins / chemistry
  • Proteins / metabolism
  • Sulfhydryl Compounds / chemistry
  • Tandem Mass Spectrometry* / methods

Substances

  • Cysteine
  • Hydroxylamine
  • Sulfhydryl Compounds
  • Proteins