Citrus Flavanone Effects on the Nrf2-Keap1/GSK3/NF-κB/NLRP3 Regulation and Corticotroph-Stress Hormone Loop in the Old Pituitary

Int J Mol Sci. 2024 Aug 16;25(16):8918. doi: 10.3390/ijms25168918.

Abstract

Oxidative stress and inflammation are significant causes of aging. At the same time, citrus flavanones, naringenin (NAR), and hesperetin (HES) are bioactives with proven antioxidant and anti-inflammatory properties. Nevertheless, there are still no data about flavanone's influence and its potential effects on the healthy aging process and improving pituitary functioning. Thus, using qPCR, immunoblot, histological techniques, and biochemical assays, our study aimed to elucidate how citrus flavanones (15 mg/kg b.m. per os) affect antioxidant defense, inflammation, and stress hormone output in the old rat model. Our results showed that HES restores the redox environment in the pituitary by down-regulating the nuclear factor erythroid 2-related factor 2 (Nrf2) protein while increasing kelch-like ECH-associated protein 1 (Keap1), thioredoxin reductase (TrxR1), and superoxide dismutase 2 (SOD2) protein expression. Immunofluorescent analysis confirmed Nrf2 and Keap1 down- and up-regulation, respectively. Supplementation with NAR increased Keap1, Trxr1, glutathione peroxidase (Gpx), and glutathione reductase (Gr) mRNA expression. Decreased oxidative stress aligned with NLRP3 decrement after both flavanones and glycogen synthase kinase-3 (GSK3) only after HES. The signal intensity of adrenocorticotropic hormone (ACTH) cells did not change, while corticosterone levels in serum decreased after both flavanones. HES showed higher potential than NAR in affecting a redox environment without increasing the inflammatory response, while a decrease in corticosterone level has a solid link to longevity. Our findings suggest that HES could improve and facilitate redox and inflammatory dysregulation in the rat's old pituitary.

Keywords: ACTH; CAT; GPx; GR; Interleukin 1 and 6; NLRP3; SOD; TNF-α; hesperetin; naringenin.

MeSH terms

  • Adrenocorticotropic Hormone / blood
  • Adrenocorticotropic Hormone / metabolism
  • Aging / drug effects
  • Aging / metabolism
  • Animals
  • Antioxidants / metabolism
  • Antioxidants / pharmacology
  • Citrus* / chemistry
  • Flavanones* / pharmacology
  • Hesperidin / pharmacology
  • Kelch-Like ECH-Associated Protein 1* / metabolism
  • Male
  • NF-E2-Related Factor 2* / metabolism
  • NF-kappa B* / metabolism
  • NLR Family, Pyrin Domain-Containing 3 Protein* / metabolism
  • Oxidative Stress / drug effects
  • Pituitary Gland* / drug effects
  • Pituitary Gland* / metabolism
  • Rats
  • Rats, Wistar
  • Signal Transduction / drug effects

Substances

  • NF-E2-Related Factor 2
  • Kelch-Like ECH-Associated Protein 1
  • Flavanones
  • NLR Family, Pyrin Domain-Containing 3 Protein
  • Nfe2l2 protein, rat
  • KEAP1 protein, rat
  • NF-kappa B
  • Adrenocorticotropic Hormone
  • Nlrp3 protein, rat
  • Antioxidants
  • hesperetin
  • naringenin
  • Hesperidin