Establishment and application of highly efficient regeneration, genetic transformation and genome editing system for cucurbitacins biosynthesis in Hemsleya chinensis

BMC Plant Biol. 2024 Nov 7;24(1):1052. doi: 10.1186/s12870-024-05717-x.

Abstract

Background: Hemsleya Chinensis is a perennial plant in the Cucurbitaceae family containing antibacterial and anti-inflammatory compounds. The lack of genetic transformation systems makes it difficult to verify the functions of genes controlling important traits and conduct molecular breeding in H. chinensis.

Results: Highly efficient calli were induced on MS medium added 1.5 mg·L- 1 6-benzylaminopurine (6-BA) and 0.02 mg·L- 1 1-naphthylacetic acid (NAA) with high efficiency (> 95%). The frequency of shoot induction was increased to 90% with a plant growth regulator combination of 1.5 mg·L- 1 6-BA and 0.1 mg·L- 1 NAA. Our results also showed that 100% of shoot regeneration was achieved in a shoot regeneration medium. Additionally, more than 92% of kanamycin-resistant plants were confirmed. Furthermore, we achieved 42% genome editing efficiency by applying this transformation method to HcOSC6, a gene that catalyzes the formation of cucurbitadienol. HPLC analysis showed OE-HcOSC6 lines exhibited significantly higher cucurbitadienol levels than the genome-edited lines. Transcriptomic analysis revealed that some downstream genes related to cucurbitadienol biosynthesis, such as HcCYP87D20, HcCYP81Q58, and HcSDR34, were up-regulated in OE lines and down-regulated in mutants.

Conclusions: Here, we established a process for regeneration, transformation, and genome editing of H. chinensis using stem segments. This provides valuable insight into the underlying molecular mechanisms of medicinal compound production. By combining high-efficiency tissue culture, transformation, and genome editing systems, we provide a powerful platform that supports functional research on molecular mechanisms of secondary metabolism.

Keywords: H. chinensis; HcOSC6; Genetic transformation system; Genome editing; Medicinal plants.

MeSH terms

  • Cucurbitaceae / genetics
  • Cucurbitaceae / metabolism
  • Cucurbitaceae / physiology
  • Gene Editing*
  • Plant Growth Regulators / metabolism
  • Plant Growth Regulators / pharmacology
  • Plant Shoots / drug effects
  • Plant Shoots / genetics
  • Plants, Genetically Modified / genetics
  • Regeneration* / drug effects
  • Regeneration* / genetics
  • Transformation, Genetic*
  • Triterpenes / metabolism

Substances

  • Triterpenes
  • Plant Growth Regulators