Abstract
The relationships between the integrin-mediated activation of inward rectifyier K+ channels (KIR), the phosphorylation of pp125FAK and the rescue of neuritogenesis were studied in 41A3 mouse neuroblastoma cells. Neuritogenesis, elicited by adhesion to FN-enriched substrata, was reversibly impaired by pretreating these cells with the tyrosine kinase inhibitor Herbimycin A. This impairment mimicked that operated by Cs+ ions, which selectively inhibited the integrin-mediated activation of KIR channels. Various phosphotyrosine containing cellular proteins underwent a marked increase upon cell adhesion to FN-coated dishes. This increase was significantly reduced by Cs+ addition. Immunoprecipitation of pp125FAK revealed that the phosphorylation of this kinase and several associated proteins was significantly and reversibly inhibited by Cs+, indicating that integrin-mediated activation of KIR channels is a limiting step upstream to the phosphorylation of pp125FAK in the commitment to neuritogenesis.
Publication types
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Research Support, Non-U.S. Gov't
MeSH terms
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Animals
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Benzoquinones
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Cell Adhesion Molecules / metabolism*
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Cell Line
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Focal Adhesion Kinase 1
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Focal Adhesion Protein-Tyrosine Kinases
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Lactams, Macrocyclic
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Membrane Potentials
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Mice
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Neoplasm Proteins / isolation & purification
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Neoplasm Proteins / metabolism
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Neurites / drug effects
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Neurites / physiology*
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Neuroblastoma
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Patch-Clamp Techniques
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Phosphoproteins / isolation & purification
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Phosphoproteins / metabolism*
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Phosphorylation
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Phosphotyrosine
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Potassium Channels / physiology*
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Potassium Channels, Inwardly Rectifying*
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Protein-Tyrosine Kinases / antagonists & inhibitors
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Protein-Tyrosine Kinases / metabolism*
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Quinones / pharmacology
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Receptor, Insulin / metabolism*
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Rifabutin / analogs & derivatives
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Tumor Cells, Cultured
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Tyrosine / analogs & derivatives
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Tyrosine / analysis
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Tyrosine / metabolism
Substances
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Benzoquinones
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Cell Adhesion Molecules
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Lactams, Macrocyclic
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Neoplasm Proteins
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Phosphoproteins
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Potassium Channels
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Potassium Channels, Inwardly Rectifying
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Quinones
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Rifabutin
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Phosphotyrosine
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Tyrosine
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herbimycin
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Protein-Tyrosine Kinases
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Receptor, Insulin
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Focal Adhesion Kinase 1
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Focal Adhesion Protein-Tyrosine Kinases
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Ptk2 protein, mouse