Elucidation of the mechanism of CryIIIA overproduction in a mutagenized strain of Bacillus thuringiensis var. tenebrionis

Mol Microbiol. 1994 Oct;14(2):381-9. doi: 10.1111/j.1365-2958.1994.tb01298.x.

Abstract

NB176 is a Bacillus thuringiensis mutant derived by gamma-irradiation of NB125 Bacillus thuringiensis var. tenebrionis (Krieg). It exhibits two interesting phenotypes: (i) oligosporogeny and (ii) twofold to threefold overproduction of the CryIIIA protein. Southern profiles of the NB176 strain showed an additional copy(s) of the cryIIIA gene located on a 4 kb HindIII fragment, in addition to the expected cryIIIA gene on a 3 kb HindIII fragment. Each cryIIIA gene-bearing HindIII fragment was cloned from NB176. The restriction map of the 3 kb HindIII fragment was identical to that published by Donovan and coworkers. Sequencing of the 4 kb HindIII fragment showed no alterations in the promoter region of the cryIIIA gene but did show replacement of the region immediately following the cryIIIA open reading frame with a sequence encoding a transposase with 50% amino acid homology to that of Tn1000. These findings suggest that the overproduction phenotype of NB176 results from extra copies of the cryIIIA gene produced from a transposition event(s) induced or stabilized by gamma-irradiation. Integration of additional copies of the cryIIIA gene into the native 90 MDa plasmid of the wild-type B. thuringiensis var. tenebrionis strain resulted in strains that made enormous crystals, many possessing greatly enhanced insecticidal activity.

MeSH terms

  • Amino Acid Sequence
  • Bacillus thuringiensis / genetics*
  • Bacillus thuringiensis / metabolism
  • Bacillus thuringiensis / radiation effects
  • Bacillus thuringiensis Toxins
  • Bacterial Proteins / biosynthesis
  • Bacterial Proteins / genetics*
  • Bacterial Toxins*
  • Cloning, Molecular
  • Endotoxins / biosynthesis
  • Endotoxins / genetics*
  • Gamma Rays
  • Gene Expression
  • Genes, Bacterial
  • Hemolysin Proteins
  • Molecular Sequence Data
  • Multigene Family
  • Mutagenesis
  • Nucleotidyltransferases / chemistry
  • Nucleotidyltransferases / genetics
  • Plasmids
  • Promoter Regions, Genetic
  • Restriction Mapping
  • Transposases

Substances

  • Bacillus thuringiensis Toxins
  • Bacterial Proteins
  • Bacterial Toxins
  • Endotoxins
  • Hemolysin Proteins
  • insecticidal crystal protein, Bacillus Thuringiensis
  • Nucleotidyltransferases
  • Transposases

Associated data

  • GENBANK/U10985