Abstract
The Syk family of protein tyrosine kinases, consisting of ZAP-70 and Syk, associate with the pre- and alphabeta T cell antigen receptors (TCRs) and undergo tyrosine phosphorylation and activation following receptor engagement. Thymocyte development in zap-70-/- mice is blocked at the CD4+CD8+ TCR(lo) stage. The presence of Syk in the thymus has raised the possibility that Syk may be able to mediate TCR function. To determine if Syk can play a role in thymocyte development, we generated zap-70-/- mice expressing a human syk cDNA. Syk expression restored both thymocyte development and function. In addition, Syk function required the CD45 transmembrane protein tyrosine phosphatase. Hence, ZAP-70 and Syk can play overlapping functions and exhibit similar regulatory mechanisms in mediating alphabeta T cell development.
Publication types
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Research Support, Non-U.S. Gov't
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Research Support, U.S. Gov't, P.H.S.
MeSH terms
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Animals
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DNA, Complementary / genetics
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DNA, Complementary / metabolism
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Enzyme Precursors / biosynthesis
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Enzyme Precursors / genetics
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Enzyme Precursors / physiology*
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Epithelium / growth & development
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Epithelium / ultrastructure
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Female
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Gene Expression
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Humans
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Intestines / growth & development
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Intestines / ultrastructure
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Intracellular Signaling Peptides and Proteins
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Leukocyte Common Antigens / genetics
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Leukocyte Common Antigens / physiology
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Mice
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Mice, Inbred C57BL
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Protein-Tyrosine Kinases / biosynthesis
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Protein-Tyrosine Kinases / genetics
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Protein-Tyrosine Kinases / physiology*
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Receptors, Antigen, T-Cell, gamma-delta / biosynthesis
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Receptors, Antigen, T-Cell, gamma-delta / physiology
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Signal Transduction / physiology
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Syk Kinase
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T-Lymphocytes / cytology
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T-Lymphocytes / physiology*
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T-Lymphocytes / ultrastructure
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Thymus Gland / cytology
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Thymus Gland / growth & development*
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Thymus Gland / ultrastructure
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Transgenes
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ZAP-70 Protein-Tyrosine Kinase
Substances
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DNA, Complementary
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Enzyme Precursors
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Intracellular Signaling Peptides and Proteins
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Receptors, Antigen, T-Cell, gamma-delta
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Protein-Tyrosine Kinases
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SYK protein, human
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Syk Kinase
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Syk protein, mouse
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ZAP-70 Protein-Tyrosine Kinase
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ZAP70 protein, human
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Zap70 protein, mouse
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Leukocyte Common Antigens