Abstract
Nuclear factor of activated T cells (NFAT) is a critical regulator of early gene transcription in response to TCR-mediated signals. Here, we show that mice lacking both NFATp and NFAT4 develop a profound lymphoproliferative disorder likely due to a lowered threshold for TCR signaling coupled with increased resistance to apoptosis secondary to defective FasL expression. NFAT mutant mice also have allergic blepharitis, interstitial pneumonitis, and a 10(3) to 10(4) fold increase in serum IgG1 and IgE levels, secondary to a dramatic and selective increase in Th2 cytokines. This phenotype may be ascribed to unopposed occupancy of the IL-4 promoter by NFATc. Our data demonstrate that lymphoid homeostasis and Th2 activation require a critical balance among NFAT family members.
Publication types
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Research Support, Non-U.S. Gov't
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Research Support, U.S. Gov't, P.H.S.
MeSH terms
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Alveolitis, Extrinsic Allergic / immunology
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Alveolitis, Extrinsic Allergic / metabolism
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Animals
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Apoptosis / physiology
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B-Lymphocytes / immunology
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B-Lymphocytes / metabolism
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B-Lymphocytes / physiology
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Blepharitis / immunology
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Blepharitis / metabolism
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Cell Nucleus / metabolism
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Cytokines / biosynthesis
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DNA-Binding Proteins / biosynthesis
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DNA-Binding Proteins / physiology*
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Fas Ligand Protein
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Homeostasis / physiology*
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Immunoglobulins / biosynthesis
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Lung Diseases, Interstitial / immunology
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Lung Diseases, Interstitial / metabolism
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Lymphocyte Activation / physiology
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Lymphoid Tissue / cytology
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Lymphoid Tissue / immunology
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Lymphoid Tissue / physiology*
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Membrane Glycoproteins / biosynthesis
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Mice
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Mice, Inbred BALB C
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NFATC Transcription Factors
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Nuclear Proteins*
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Receptors, Antigen, T-Cell / physiology
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Signal Transduction / physiology
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T-Lymphocytes / immunology
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T-Lymphocytes / metabolism
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T-Lymphocytes / physiology
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Th2 Cells / immunology
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Th2 Cells / metabolism
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Th2 Cells / physiology*
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Transcription Factors / biosynthesis
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Transcription Factors / physiology*
Substances
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Cytokines
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DNA-Binding Proteins
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Fas Ligand Protein
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Fasl protein, mouse
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Immunoglobulins
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Membrane Glycoproteins
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NFATC Transcription Factors
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Nfatc3 protein, mouse
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Nuclear Proteins
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Receptors, Antigen, T-Cell
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Transcription Factors