Princess Amberle Elessedil is selected to be a member of the Chosen that cares for the most sacred Elven tree, the Ellcrys. However she is given a vision of the Ellcrys dying, which could mean trouble for the Four Lands.
The Dagda Mor sends the Reaper after Amberle and Wil as they are taken prisoner, once again, by Cephelo. Allanon trains Bandon. Ander is forced to make a difficult decision.
To throw the demons off their trail, Amberle, Wil, and company take shelter in the abandoned fortress of Pykon. But is it abandoned? The Changeling twists Arion against Allanon.
Having been separated after their fall, Wil meets up with a Winged-Elf named Perk while Amberle and Eretria are on the run from elf hunters. Eventine sends Ander and Arion to kill the Dagda Mor. Allanon tries to heal his wounds.
Amberle, Wil, and Eretria work their way through the tunnels of Safehold. Allanon tries to help Bandon fight the Dagda Mor's will. Ander finds his rule tested.
Wil and Amberle race to reach the Ellcrys while Allanon and Ander lead the Elvin army in an epic battle against the Dagda Mor and his Demon horde. But the cost of their quest could prove too great.