Abstract
In the chick embryo, left-right asymmetric patterns of gene expression in the lateral plate mesoderm are initiated by signals located in and around Hensen's node. Here we show that Caronte (Car), a secreted protein encoded by a member of the Cerberus/Dan gene family, mediates the Sonic hedgehog (Shh)-dependent induction of left-specific genes in the lateral plate mesoderm. Car is induced by Shh and repressed by fibroblast growth factor-8 (FGF-8). Car activates the expression of Nodal by antagonizing a repressive activity of bone morphogenic proteins (BMPs). Our results define a complex network of antagonistic molecular interactions between Activin, FGF-8, Lefty-1, Nodal, BMPs and Car that cooperate to control left-right asymmetry in the chick embryo.
Publication types
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Research Support, Non-U.S. Gov't
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Research Support, U.S. Gov't, P.H.S.
MeSH terms
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Amino Acid Sequence
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Animals
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Avian Proteins*
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Body Patterning / genetics
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Body Patterning / physiology*
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Bone Morphogenetic Proteins / physiology
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COS Cells
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Chick Embryo
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Cloning, Molecular
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Fibroblast Growth Factor 8
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Fibroblast Growth Factors / physiology
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Gene Expression Regulation, Developmental
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Genes, Homeobox
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Hedgehog Proteins
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Homeodomain Proteins / physiology
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Humans
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Intercellular Signaling Peptides and Proteins
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Left-Right Determination Factors
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Molecular Sequence Data
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Nodal Protein
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Proteins / genetics
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Proteins / physiology*
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Sequence Homology, Amino Acid
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Trans-Activators*
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Transcription Factors / physiology
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Transforming Growth Factor beta / physiology
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Xenopus Proteins
Substances
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Avian Proteins
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Bone Morphogenetic Proteins
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FGF8 protein, human
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Hedgehog Proteins
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Homeodomain Proteins
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Intercellular Signaling Peptides and Proteins
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LEFTY1 protein, human
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Left-Right Determination Factors
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NKX3-2 protein, human
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NODAL protein, human
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Nodal Protein
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Proteins
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SHH protein, human
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Trans-Activators
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Transcription Factors
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Transforming Growth Factor beta
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Xenopus Proteins
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caronte protein, Gallus gallus
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cer1 protein, Xenopus
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Fibroblast Growth Factor 8
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Fibroblast Growth Factors