Abstract
We recently reported that the number of gamma delta T cells was increased after infection with Escherichia coli in C3H/HeN mice. We here showed that an i.p. injection with native lipid A derived from E. coli induced an increase of gamma delta T cells in the peritoneal cavity of LPS-responsive C3H/HeN mice and, albeit to a lesser degree, also in LPS-hyporesponsive C3H/HeJ mice. The purified gamma delta T cells from C3H/HeN and C3H/HeJ mice expressed a canonical TCR repertoire encoded by V gamma 6-J gamma 1/V delta 1-D delta 2-J delta 2 gene segments and proliferated in response to the native lipid A derived from E. coli in a TCR-independent manner. The lipid A-reactive gamma delta T cells bearing canonical V gamma 6/V delta 1 expressed Toll-like receptor (TLR) 2 mRNA, while TLR4 mRNA was undetectable. Treatment with a TLR2 anti-sense oligonucleotide resulted in hyporesponsiveness of the gamma delta T cells to the native lipid A. TLR2-deficient mice showed an impaired increase of the gamma delta T cells following injection of native lipid A. These results suggest that TLR2 is involved in the activation of canonical V gamma 6/V delta 1 T cells by native E. coli lipid A.
Publication types
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Research Support, Non-U.S. Gov't
MeSH terms
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Animals
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Ascitic Fluid / immunology
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Ascitic Fluid / metabolism
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Cell Line
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Cytokines / biosynthesis
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Drosophila Proteins*
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Escherichia coli Infections / immunology*
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Escherichia coli Infections / metabolism
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Female
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Gene Expression Regulation / immunology
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Genes, T-Cell Receptor delta
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Genes, T-Cell Receptor gamma
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Injections, Intraperitoneal
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Lipid A / administration & dosage
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Lipid A / physiology
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Lipopolysaccharide Receptors / biosynthesis
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Lipopolysaccharide Receptors / metabolism
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Lipopolysaccharides / metabolism
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Lipopolysaccharides / pharmacology
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Lymphocyte Activation
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Membrane Glycoproteins / biosynthesis*
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Membrane Glycoproteins / genetics
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Membrane Glycoproteins / physiology
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Mice
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Mice, Inbred C3H
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Mice, Inbred C57BL
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Mice, Knockout
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Peritoneal Cavity / cytology
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RNA, Messenger / biosynthesis
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Receptors, Antigen, T-Cell, gamma-delta / biosynthesis*
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Receptors, Antigen, T-Cell, gamma-delta / genetics
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Receptors, Cell Surface / biosynthesis*
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Receptors, Cell Surface / genetics
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Receptors, Cell Surface / physiology
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T-Lymphocyte Subsets / metabolism*
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Toll-Like Receptor 2
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Toll-Like Receptor 4
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Toll-Like Receptors
Substances
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Cytokines
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Drosophila Proteins
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Lipid A
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Lipopolysaccharide Receptors
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Lipopolysaccharides
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Membrane Glycoproteins
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RNA, Messenger
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Receptors, Antigen, T-Cell, gamma-delta
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Receptors, Cell Surface
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Toll-Like Receptor 2
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Toll-Like Receptor 4
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Toll-Like Receptors