neuralized functions cell-autonomously to regulate a subset of notch-dependent processes during adult Drosophila development

Dev Biol. 2001 Mar 1;231(1):217-33. doi: 10.1006/dbio.2000.0124.

Abstract

neuralized (neu) represents one of the strong neurogenic mutants in Drosophila. Mutants of this class display, among other phenotypes, a strong overcommitment to neural fates at the expense of epidermal fates. We analyzed the role of neu during adult development by using mutant clonal analysis, misexpression of wild-type and truncated forms of Neu, and examination of genetic interactions with N-pathway mutations. We find that neu is required cell-autonomously for lateral inhibition during peripheral neurogenesis and for multiple asymmetric cell divisions in the sensory lineage. In contrast, neu is apparently dispensable for other N-mediated processes, including lateral inhibition during wing vein development and wing margin induction. Misexpression of wild-type Neu causes defects in both peripheral neurogenesis and wing vein development, while a truncated form lacking the RING finger is further capable of inhibiting formation of the wing margin. In addition, the phenotypes produced by misexpression of wild-type and truncated Neu proteins are sensitive to the dosage of several N-pathway components. Finally, using epitope-tagged Neu proteins, we localize Neu to the plasma membrane and reveal a novel morphology to the sensory organ precursor cells of wing imaginal discs. Collectively, these data indicate a key role for neu in the reception of the lateral inhibitory signal during peripheral neurogenesis.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Animals
  • Drosophila / growth & development*
  • Drosophila Proteins*
  • Larva / physiology
  • Ligases*
  • Membrane Proteins / physiology*
  • Nerve Tissue Proteins / physiology*
  • Peripheral Nerves / growth & development
  • Receptors, Notch
  • Ubiquitin-Protein Ligases*
  • Wings, Animal / growth & development

Substances

  • Drosophila Proteins
  • Membrane Proteins
  • N protein, Drosophila
  • Nerve Tissue Proteins
  • Receptors, Notch
  • Ubiquitin-Protein Ligases
  • neur protein, Drosophila
  • Ligases