Abstract
The transcriptional coactivator CREB-binding protein (CBP) is known to play an important role in coupling signal transduction pathways to changes in gene expression. In many cases, this is achieved by the stimulus-specific recruitment of CBP to promoter-bound transcription factors. However, a number of recent studies have suggested that signal transduction pathways can also directly influence CBP-mediated transcriptional activity. Here we show that in Jurkat cells the activity of the CBP C-terminal transactivation domain is strongly upregulated in response to either T cell receptor stimulation or the combination of ionomycin and phorbol ester. We further show that maximal stimulation of CBP-mediated transcription requires the synergistic activation of both the calcineurin and Ras-MAPK signaling pathways. These results indicate that CBP can function as a T cell activation-inducible transcriptional coactivator and is therefore likely to play an important role in T cell activation-induced gene expression.
Copyright 2001 Academic Press.
Publication types
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Research Support, U.S. Gov't, P.H.S.
MeSH terms
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Antibodies, Monoclonal / pharmacology
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CD3 Complex / immunology
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CREB-Binding Protein
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Calcineurin / metabolism
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Calcineurin Inhibitors
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Calcium-Calmodulin-Dependent Protein Kinases / antagonists & inhibitors
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Calcium-Calmodulin-Dependent Protein Kinases / metabolism
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DNA, Recombinant
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Enzyme Inhibitors / pharmacology
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Flavonoids / pharmacology
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Gene Expression Regulation / drug effects
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Humans
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Ionomycin / pharmacology
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Ionophores / pharmacology
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Jurkat Cells
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Luciferases / genetics
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Luciferases / metabolism
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Nuclear Proteins / genetics
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Nuclear Proteins / physiology*
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Plasmids / genetics
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Recombinant Fusion Proteins / genetics
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Recombinant Fusion Proteins / metabolism
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Signal Transduction / drug effects
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T-Lymphocytes / cytology
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T-Lymphocytes / metabolism*
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Tacrolimus / pharmacology
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Tetradecanoylphorbol Acetate / pharmacology
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Trans-Activators / genetics
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Trans-Activators / physiology*
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Transcription, Genetic
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Transfection
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Up-Regulation
Substances
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Antibodies, Monoclonal
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CD3 Complex
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Calcineurin Inhibitors
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DNA, Recombinant
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Enzyme Inhibitors
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Flavonoids
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Ionophores
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Nuclear Proteins
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Recombinant Fusion Proteins
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Trans-Activators
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Ionomycin
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Luciferases
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CREB-Binding Protein
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CREBBP protein, human
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Calcium-Calmodulin-Dependent Protein Kinases
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Calcineurin
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Tetradecanoylphorbol Acetate
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2-(2-amino-3-methoxyphenyl)-4H-1-benzopyran-4-one
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Tacrolimus