Abstract
TRAIL (tumour-necrosis factor-related apoptosis ligand or Apo2L) triggers apoptosis through engagement of the death receptors TRAIL-R1 (also known as DR4) and TRAIL-R2 (DR5). Here we show that the c-Rel subunit of the transcription factor NF-kappaB induces expression of TRAIL-R1 and TRAIL-R2; conversely, a transdominant mutant of the inhibitory protein IkappaBalpha or a transactivation-deficient mutant of c-Rel reduces expression of either death receptor. Whereas NF-kappaB promotes death receptor expression, cytokine-mediated activation of the RelA subunit of NF-kappaB also increases expression of the apoptosis inhibitor, Bcl-xL, and protects cells from TRAIL. Inhibition of NF-kappaB by blocking activation of the IkappaB kinase complex reduces Bcl-x L expression and sensitizes tumour cells to TRAIL-induced apoptosis. The ability to induce death receptors or Bcl-xL may explain the dual roles of NF-kappaB as a mediator or inhibitor of cell death during immune and stress responses.
Publication types
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Research Support, Non-U.S. Gov't
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Research Support, U.S. Gov't, Non-P.H.S.
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Research Support, U.S. Gov't, P.H.S.
MeSH terms
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Animals
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Apoptosis
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Apoptosis Regulatory Proteins
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DNA-Binding Proteins / metabolism
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Gene Expression Regulation*
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HeLa Cells
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Humans
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I-kappa B Proteins*
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Membrane Glycoproteins / metabolism*
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Membrane Glycoproteins / pharmacology
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Mice
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NF-KappaB Inhibitor alpha
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NF-kappa B / genetics
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NF-kappa B / metabolism*
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Proto-Oncogene Proteins c-bcl-2 / metabolism
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Proto-Oncogene Proteins c-rel / genetics
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Proto-Oncogene Proteins c-rel / metabolism*
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Radiation Tolerance
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Receptors, TNF-Related Apoptosis-Inducing Ligand
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Receptors, Tumor Necrosis Factor / biosynthesis
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Receptors, Tumor Necrosis Factor / genetics*
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TNF-Related Apoptosis-Inducing Ligand
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Transcription Factor RelA
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Tumor Cells, Cultured
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Tumor Necrosis Factor-alpha / metabolism*
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Tumor Necrosis Factor-alpha / pharmacology
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Tumor Suppressor Protein p53 / genetics
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Tumor Suppressor Protein p53 / metabolism
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bcl-X Protein
Substances
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Apoptosis Regulatory Proteins
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BCL2L1 protein, human
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Bcl2l1 protein, mouse
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DNA-Binding Proteins
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I-kappa B Proteins
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Membrane Glycoproteins
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NF-kappa B
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NFKBIA protein, human
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Nfkbia protein, mouse
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Proto-Oncogene Proteins c-bcl-2
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Proto-Oncogene Proteins c-rel
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Receptors, TNF-Related Apoptosis-Inducing Ligand
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Receptors, Tumor Necrosis Factor
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TNF-Related Apoptosis-Inducing Ligand
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TNFRSF10A protein, human
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TNFRSF10B protein, human
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TNFSF10 protein, human
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Tnfrsf10b protein, mouse
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Tnfsf10 protein, mouse
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Transcription Factor RelA
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Tumor Necrosis Factor-alpha
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Tumor Suppressor Protein p53
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bcl-X Protein
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NF-KappaB Inhibitor alpha