The aerobic physiology of the air-breathing blue gourami, Trichogaster trichopterus, necessitates behavioural regulation of breath-hold limits during hypoxic stress and predatory challenge

J Comp Physiol B. 2001 Oct;171(7):603-12. doi: 10.1007/s003600100211.

Abstract

Physiological characteristics of the blood oxygen transport system and muscle metabolism indicate a high dependence on aerobic pathways in the blue gourami, Trichogaster trichopterus. Haemoglobin concentration and haematocrit were modest and the blood oxygen affinity (P50=2.31 kPa at pH 7.4 and 28 degrees C) and its sensitivity to pH (Bohr factor, phi=-0.34) favour oxygen unloading at a relatively high oxygen pressure (PO2). The intracellular buffering capacity (44.0 slykes) and lactate dehydrogenase (LDH) activity (154.3 iu g(-1)) do not support exceptional anaerobic capabilities. Air-breathing frequency in the blue gourami is expected to increase when aquatic oxygen tensions decline. Under threat of predation, however, this behaviour must be modified at a potential cost to aerobic metabolism. We therefore tested the hypothesis that metabolic responses to predatory challenge and aquatic hypoxia are subject to behavioural modulation. Computer-generated visual stimuli consistently reduced air-breathing frequency at 19.95, 6.65 and 3.33 kPa PO2. Bi-directional rates of spontaneous activity were similarly reduced. The metabolic cost of this behaviour was estimated and positively correlated with PO2 but not with visual stimulation thus indicating down-regulation of spontaneous activity rather than breath-holding behaviour. Neither PO2 nor visual stimulation resulted in significant change to muscle lactate and ATP concentrations and confirm that aerobic breath-hold limits were maintained following behavioural modulation of metabolic demands.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Adaptation, Physiological / physiology*
  • Adenosine Diphosphate / blood
  • Adenosine Triphosphate / blood
  • Air
  • Animals
  • Environment Design
  • Fishes / physiology*
  • Guanosine Triphosphate / blood
  • Hemoglobins
  • Hypoxia / metabolism
  • Hypoxia / physiopathology*
  • L-Lactate Dehydrogenase / metabolism
  • Lactates / metabolism
  • Muscle, Skeletal / metabolism
  • Oxygen / blood
  • Oxyhemoglobins / metabolism
  • Predatory Behavior
  • Respiration
  • Stress, Physiological / metabolism
  • Stress, Physiological / physiopathology*

Substances

  • Hemoglobins
  • Lactates
  • Oxyhemoglobins
  • Adenosine Diphosphate
  • Guanosine Triphosphate
  • Adenosine Triphosphate
  • L-Lactate Dehydrogenase
  • Oxygen