Abstract
Gemin3 is a DEAD-box RNA helicase that binds to the Survival of Motor Neurons (SMN) protein and is a component of the SMN complex, which also comprises SMN, Gemin2, Gemin4, Gemin5, and Gemin6. Reduction in SMN protein results in Spinal muscular atrophy (SMA), a common neurodegenerative disease. The SMN complex has critical functions in the assembly/restructuring of diverse ribonucleoprotein (RNP) complexes. Here we report that Gemin3 and Gemin4 are also in a separate complex that contains eIF2C2, a member of the Argonaute protein family. This novel complex is a large approximately 15S RNP that contains numerous microRNAs (miRNAs). We describe 40 miRNAs, a few of which are identical to recently described human miRNAs, a class of small endogenous RNAs. The genomic sequences predict that miRNAs are likely to be derived from larger precursors that have the capacity to form stem-loop structures.
Publication types
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Research Support, Non-U.S. Gov't
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Research Support, U.S. Gov't, P.H.S.
MeSH terms
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Animals
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Argonaute Proteins
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Blotting, Western
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Centrifugation, Density Gradient
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Cloning, Molecular
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DEAD Box Protein 20
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DEAD-box RNA Helicases
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Eukaryotic Initiation Factors*
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HeLa Cells
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Humans
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Mice
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MicroRNAs
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Minor Histocompatibility Antigens
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Nerve Tissue Proteins / metabolism
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Nuclear Proteins / metabolism
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Nucleic Acid Conformation
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Peptide Initiation Factors / metabolism
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Plasmids / metabolism
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Precipitin Tests
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Protein Binding
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Protein Structure, Tertiary
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RNA Helicases / metabolism
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RNA, Antisense / chemistry*
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RNA, Antisense / classification
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RNA, Messenger / metabolism
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RNA, Untranslated / chemistry*
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RNA, Untranslated / classification
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RNA-Binding Proteins
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Ribonucleoproteins / chemistry*
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Ribonucleoproteins / metabolism*
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Ribonucleoproteins, Small Nuclear
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SMN Complex Proteins
Substances
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AGO1 protein, human
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Argonaute Proteins
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Eukaryotic Initiation Factors
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GEMIN2 protein, human
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GEMIN4 protein, human
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GEMIN5 protein, human
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GEMIN6 protein, human
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Gemin5 protein, mouse
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Gemin6 protein, mouse
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MicroRNAs
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Minor Histocompatibility Antigens
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Nerve Tissue Proteins
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Nuclear Proteins
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Peptide Initiation Factors
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RNA, Antisense
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RNA, Messenger
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RNA, Untranslated
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RNA-Binding Proteins
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Ribonucleoproteins
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Ribonucleoproteins, Small Nuclear
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SMN Complex Proteins
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Sip1 protein, mouse
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DDX20 protein, human
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DEAD Box Protein 20
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Ddx20 protein, mouse
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DEAD-box RNA Helicases
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RNA Helicases