Abstract
Endocytosed microbial antigens are primarily delivered to lysosomal compartments where antigen binding to MHC and CD1 molecules occurs in an acidic and proteolytically active environment. Signal-dependent delivery to lysosomes has been suggested for these antigen-presenting molecules, but molecular interactions with vesicular coat proteins and adaptors that direct their lysosomal sorting are poorly understood. Here CD1b but not other CD1 isoforms bound the AP-3 adaptor protein complex. In AP-3-deficient cells derived from patients with Hermansky-Pudlak syndrome type 2 (HPS-2), CD1b failed to efficiently gain access to lysosomes, resulting in a profound defect in antigen presentation. Since MHC class II traffics normally in AP-3-deficient cells, defects in CD1b antigen presentation may account for recurrent bacterial infections in HPS-2 patients.
Publication types
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Research Support, Non-U.S. Gov't
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Research Support, U.S. Gov't, P.H.S.
MeSH terms
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Adaptor Proteins, Vesicular Transport
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Amino Acid Motifs
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Antigen Presentation*
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Antigens, Bacterial / immunology
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Antigens, CD / chemistry
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Antigens, CD / metabolism
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Antigens, CD1 / chemistry
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Antigens, CD1 / metabolism*
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Antigens, CD1 / physiology*
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Bacterial Infections / immunology
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Carrier Proteins / genetics*
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Carrier Proteins / metabolism
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Cell Line
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Gene Deletion
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Glycoproteins / chemistry
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Glycoproteins / metabolism
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HeLa Cells
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Humans
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Lipids / immunology
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Lysosomes / metabolism
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Lysosomes / ultrastructure
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Macromolecular Substances
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Membrane Proteins / genetics*
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Membrane Proteins / metabolism
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Monomeric Clathrin Assembly Proteins*
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Platelet Membrane Glycoproteins / chemistry
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Platelet Membrane Glycoproteins / metabolism
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Protein Isoforms / metabolism
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Protein Transport
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Syndrome
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Tetraspanin 30
Substances
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Adaptor Proteins, Vesicular Transport
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Antigens, Bacterial
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Antigens, CD
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Antigens, CD1
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CD1C protein, human
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CD1b antigen
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CD63 protein, human
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Carrier Proteins
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Glycoproteins
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Lipids
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Macromolecular Substances
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Membrane Proteins
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Monomeric Clathrin Assembly Proteins
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Platelet Membrane Glycoproteins
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Protein Isoforms
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Tetraspanin 30
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clathrin assembly protein AP180