Abstract
UFD2a is a mammalian homolog of Saccharomyces cerevisiae Ufd2, originally described as an E4 ubiquitination factor. UFD2a belongs to the U-box family of ubiquitin ligases (E3s) and likely functions as both an E3 and E4. We have isolated and characterized the mouse gene (Ube4b) for UFD2a. A full-length (approximately 5700 bp) Ube4b cDNA was isolated and the corresponding gene spans >100 kb, comprising 27 exons. Luciferase reporter gene analysis of the 5(') flanking region of Ube4b revealed that nucleotides -1018 to -943 (relative to the translation initiation site) possess promoter activity. This functional sequence contains two putative Sp1 binding sites but not a TATA box. Immunoblot and immunohistochemical analyses revealed that UFD2a is expressed predominantly in the neuronal tissues. We also show that UFD2a interacts with VCP (a AAA-family ATPase) that is thought to mediate protein folding. These data implicate UFD2a in the degradation of neuronal proteins by the ubiquitin-proteasome pathway.
Publication types
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Research Support, Non-U.S. Gov't
MeSH terms
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Adenosine Triphosphatases
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Animals
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Base Sequence
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Cell Cycle Proteins / metabolism
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DNA, Complementary / isolation & purification
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Fungal Proteins / genetics
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Gene Components
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Ligases / genetics*
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Ligases / metabolism
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Mice
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Molecular Sequence Data
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Neurons / metabolism
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Promoter Regions, Genetic
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Saccharomyces cerevisiae Proteins*
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Transcriptional Activation
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Ubiquitin-Conjugating Enzymes
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Ubiquitin-Protein Ligases
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Valosin Containing Protein
Substances
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Cell Cycle Proteins
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DNA, Complementary
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Fungal Proteins
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Saccharomyces cerevisiae Proteins
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Ubiquitin-Conjugating Enzymes
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Ube4b protein, mouse
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Ubiquitin-Protein Ligases
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Adenosine Triphosphatases
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CDC48 protein, S cerevisiae
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Valosin Containing Protein
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Vcp protein, mouse
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Ligases
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UFD2 protein, S cerevisiae
Associated data
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GENBANK/AB083248
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GENBANK/AB083249
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GENBANK/AB083250
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GENBANK/AB083251
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GENBANK/AB083252
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GENBANK/AB083253
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GENBANK/AB083254
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GENBANK/AB083255
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GENBANK/AB083256
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GENBANK/AB083257
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GENBANK/AB083258
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GENBANK/AB083259
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GENBANK/AB083260
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GENBANK/AB083261
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GENBANK/AB083262
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GENBANK/AB083263
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GENBANK/AB083264
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GENBANK/AB083265
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GENBANK/AB083266
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GENBANK/AB083267
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GENBANK/AB083268
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GENBANK/AB083269
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GENBANK/AB083270
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GENBANK/AB083271
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GENBANK/AB083272
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GENBANK/AB083273
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GENBANK/AB083274