Cutting edge: efficient MHC class I cross-presentation during early vaccinia infection requires the transfer of proteasomal intermediates between antigen donor and presenting cells

J Immunol. 2003 Dec 1;171(11):5668-72. doi: 10.4049/jimmunol.171.11.5668.

Abstract

Priming of CD8(+) T cells requires presentation of short peptides bound to MHC class I molecules of professional APCs. Cross-presentation is a mechanism whereby professional APC present on their own MHC class I molecules peptides derived from degradation of Ags synthesized by other Ag "donor cells." The mechanism of cross-presentation is poorly understood, and the nature of the transferred Ag is unknown. In this report, we demonstrate that the bulk of a cross-presented Ag transferred from donor cells recently infected with vaccinia virus are proteasomal products that are susceptible to peptidases within the donor cell cytosol and not full-length proteins or mature epitopes either free or bound to chaperones.

Publication types

  • Research Support, U.S. Gov't, P.H.S.

MeSH terms

  • Animals
  • Antigen Presentation / immunology*
  • Antigen-Presenting Cells / immunology*
  • Antigen-Presenting Cells / metabolism*
  • Antigen-Presenting Cells / virology
  • Antigens / metabolism*
  • Cell Line
  • Chickens
  • Cysteine Endopeptidases / metabolism*
  • Cytosol / immunology
  • Cytosol / metabolism
  • Egg Proteins / immunology
  • Egg Proteins / metabolism
  • Endoplasmic Reticulum / immunology
  • Endoplasmic Reticulum / metabolism
  • H-2 Antigens / immunology
  • H-2 Antigens / metabolism*
  • HeLa Cells
  • Humans
  • Hybridomas
  • Immunodominant Epitopes / immunology
  • Immunodominant Epitopes / metabolism
  • Mice
  • Molecular Chaperones / metabolism
  • Multienzyme Complexes / metabolism*
  • Ovalbumin / immunology
  • Ovalbumin / metabolism
  • Peptide Fragments
  • Proteasome Endopeptidase Complex
  • Protein Binding / immunology
  • Protein Transport / immunology
  • Vaccinia virus / immunology*

Substances

  • Antigens
  • Egg Proteins
  • H-2 Antigens
  • Immunodominant Epitopes
  • Molecular Chaperones
  • Multienzyme Complexes
  • OVA-8
  • Peptide Fragments
  • Ovalbumin
  • Cysteine Endopeptidases
  • Proteasome Endopeptidase Complex