Nitric oxide generated from isoniazid activation by KatG: source of nitric oxide and activity against Mycobacterium tuberculosis

Antimicrob Agents Chemother. 2004 Aug;48(8):3006-9. doi: 10.1128/AAC.48.8.3006-3009.2004.

Abstract

Isonicotinic acid hydrazide (INH) is a frontline antituberculosis agent. Once taken up by Mycobacterium tuberculosis, INH requires activation by the catalase-peroxidase KatG, converting INH from its prodrug form into a range of bactericidal reactive species. Here we used 15N-labeled INH together with electron paramagnetic resonance spin trapping techniques to demonstrate that nitric oxide (NO*) is generated from oxidation at the hydrazide nitrogens during the activation of INH by M. tuberculosis KatG. We also observed that a specific scavenger of NO* provided protection against the antimycobacterial activity of INH in bacterial culture. No significant increases in mycobacterial protein nitration were detected, suggesting that NOdot; and not peroxynitrite, a nitrating metabolite of NO*, is involved in antimycobacterial action. In conclusion, INH-derived NO* has biological activity, which directly contributes to the antimycobacterial action of INH.

Publication types

  • Research Support, U.S. Gov't, P.H.S.

MeSH terms

  • Antitubercular Agents / metabolism
  • Antitubercular Agents / pharmacology*
  • Bacterial Proteins*
  • Biotransformation
  • Catalase / metabolism
  • Free Radical Scavengers / pharmacology
  • Isoniazid / metabolism
  • Isoniazid / pharmacology*
  • Mycobacterium bovis / metabolism
  • Mycobacterium tuberculosis / drug effects*
  • Mycobacterium tuberculosis / genetics
  • Mycobacterium tuberculosis / metabolism
  • Nitric Oxide / metabolism*
  • Oxidoreductases / genetics
  • Oxidoreductases / metabolism
  • Oxidoreductases / physiology*
  • Peroxidase / metabolism
  • Peroxynitrous Acid / metabolism
  • Prodrugs / metabolism
  • Prodrugs / pharmacology
  • Spin Trapping
  • Tyrosine / analogs & derivatives*
  • Tyrosine / metabolism

Substances

  • Antitubercular Agents
  • Bacterial Proteins
  • Free Radical Scavengers
  • Prodrugs
  • Peroxynitrous Acid
  • Nitric Oxide
  • 3-nitrotyrosine
  • Tyrosine
  • Oxidoreductases
  • Catalase
  • katG protein, Mycobacterium tuberculosis
  • Peroxidase
  • Isoniazid