Nucleostemin, a coordinator of self-renewal, is expressed in rat marrow stromal cells and turns off after induction of neural differentiation

Neurosci Lett. 2005 Dec 23;390(2):81-6. doi: 10.1016/j.neulet.2005.08.011. Epub 2005 Sep 1.

Abstract

Bone marrow stromal cells (BMSCs) are pluripotent stem cells with self-renewal property and potential to differentiate into a variety of cell types. Identification of the genes responsible for coordination of these processes and elucidation of the mechanisms underlying these events are of fundamental importance. Nucleostemin, a novel p53 binding protein localized in the nucleoli of ESCs, is not expressed in the differentiated cells of adult tissue, suggesting a role in maintaining stem cell self-renewal. In the present study, we have evaluated the expression profile of nucleostemin in rat BMSCs before and after the induction of neural differentiation by RT-PCR and immunocytochemistry. The profile of nucleostemin expression is then compared to the key regulators of proliferation/differentiation such as: Oct-4, Nanog, Neuro D, and Cyclin D1. Our data reveal that there is no detectable expression of Oct-4 and Nanog in either non-differentiated or neurally differentiated BMSCs. In contrast, the expression of nucleostemin is relatively high in non-differentiated BMSCs, then sharply diminishes upon induction of differentiation and finally completely vanishes by 6h after initiation of differentiation. The disappearance of nucleostemin expression coincides with the appearance of Neurofilament-M and -H, MAP2, synaptophysin- and neuron-specific enolase as revealed by RT-PCR and/or immunocytochemistry. Expression of Neuro D and Cyclin D1 also diminish as differentiation proceeds but at much slower rates as compared to nucleostemin. In conclusion, our results suggest that nucleostemin, but not Oct-4 or Nanog, is expressed in BMSCs and it possibly regulates self-renewal proliferation in BMSCs.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Animals
  • Biomarkers / metabolism
  • Bone Marrow Cells / cytology*
  • Bone Marrow Cells / metabolism
  • Carrier Proteins
  • Cell Differentiation / physiology*
  • Cell Shape
  • DNA-Binding Proteins / metabolism
  • GTP-Binding Proteins
  • Gene Expression Regulation
  • Homeodomain Proteins / metabolism
  • Neurons / cytology
  • Neurons / physiology*
  • Nuclear Proteins
  • Octamer Transcription Factor-3 / metabolism
  • Rats
  • Stromal Cells / cytology
  • Stromal Cells / physiology*

Substances

  • Biomarkers
  • Carrier Proteins
  • DNA-Binding Proteins
  • Gnl3 protein, rat
  • Homeodomain Proteins
  • Nuclear Proteins
  • Octamer Transcription Factor-3
  • GTP-Binding Proteins