Quantification of isozyme-specific activation of phospholipase C-beta2 by Rac GTPases and phospholipase C-epsilon by Rho GTPases in an intact cell assay system

Methods Enzymol. 2006:406:489-99. doi: 10.1016/S0076-6879(06)06037-X.

Abstract

Phospholipase C (PLC) catalyzes the hydrolysis of PtdIns(4,5)P2, which results in both formation of the second messengers Ins(1,4,5)P3 and diacylglycerol and alteration in the membrane association and/or activity of PtdIns(4,5)P2-binding proteins. The existence of 13 different PLC isozymes suggests multiple mechanisms of regulation of inositol lipid signaling, and the recent realization that Rho-family GTPases directly bind and activate certain PLC isozymes has added to this potential diversity of inositol lipid-related signal transduction. With the goal of delineating a less labor-intensive method for quantification of intracellular inositol phosphate production, we have applied a commercially available yttrium silicate RNA binding resin selective for inositol phosphates to develop a high-throughput inositol phosphate scintillation proximity assay (SPA). We highlight the utility of this assay using COS-7 cells robotically transfected in a 96-well format. This method is readily applied to quantify activation of PLC by receptors and G proteins, and we illustrate here the selective activation of PLC-beta2 by Rac but not by Rho GTPases and the selective activation of PLC-epsilon by Rho but not Rac GTPases.

Publication types

  • Research Support, N.I.H., Extramural

MeSH terms

  • Animals
  • COS Cells
  • Chlorocebus aethiops
  • Enzyme Activation
  • Isoenzymes / metabolism*
  • Phosphatidylinositols / metabolism
  • Phosphoinositide Phospholipase C
  • Phospholipase C beta
  • Transfection / methods
  • Type C Phospholipases / metabolism*
  • rac GTP-Binding Proteins / metabolism*
  • rho GTP-Binding Proteins / metabolism*

Substances

  • Isoenzymes
  • Phosphatidylinositols
  • Type C Phospholipases
  • Phosphoinositide Phospholipase C
  • Phospholipase C beta
  • phospholipase C epsilon
  • rac GTP-Binding Proteins
  • rho GTP-Binding Proteins