Converting IL-15 to a superagonist by binding to soluble IL-15R{alpha}

Proc Natl Acad Sci U S A. 2006 Jun 13;103(24):9166-71. doi: 10.1073/pnas.0600240103. Epub 2006 Jun 6.

Abstract

IL-15 is normally presented in vivo as a cell-associated cytokine bound to IL-15Ralpha. We show here that the biological activity of soluble IL-15 is much improved after interaction with recombinant soluble IL-15Ralpha; after injection, soluble IL-15/IL-15Ralpha complexes rapidly induce strong and selective expansion of memory-phenotype CD8(+) cells and natural killer cells. These findings imply that binding of IL-15Ralpha to IL-15 may create a conformational change that potentiates IL-15 recognition by the betagamma(c) receptor on T cells. The enhancing effect of IL-15Ralpha binding may explain why IL-15 normally functions as a cell-associated cytokine. Significantly, the results with IL-2, a soluble cytokine, are quite different; thus, IL-2 function is markedly inhibited by binding to soluble IL-2Ralpha.

Publication types

  • Research Support, N.I.H., Extramural
  • Research Support, Non-U.S. Gov't
  • Research Support, U.S. Gov't, Non-P.H.S.

MeSH terms

  • Animals
  • CD8-Positive T-Lymphocytes / metabolism
  • Cell Proliferation
  • Humans
  • Interleukin-15 / genetics
  • Interleukin-15 / metabolism*
  • Interleukin-2 / genetics
  • Interleukin-2 / metabolism
  • Interleukin-2 Receptor alpha Subunit
  • Mice
  • Mice, Inbred C57BL
  • Mice, Knockout
  • Protein Binding
  • Protein Isoforms / genetics
  • Protein Isoforms / metabolism*
  • Receptors, Interleukin-15
  • Receptors, Interleukin-2 / genetics
  • Receptors, Interleukin-2 / metabolism*
  • Recombinant Fusion Proteins / genetics
  • Recombinant Fusion Proteins / metabolism
  • Recombinant Proteins / genetics
  • Recombinant Proteins / metabolism

Substances

  • IL15RA protein, human
  • IL2RA protein, human
  • Il15ra protein, mouse
  • Il2ra protein, mouse
  • Interleukin-15
  • Interleukin-2
  • Interleukin-2 Receptor alpha Subunit
  • Protein Isoforms
  • Receptors, Interleukin-15
  • Receptors, Interleukin-2
  • Recombinant Fusion Proteins
  • Recombinant Proteins