Sequential processing of a mitochondrial tandem protein: insights into protein import in Schizosaccharomyces pombe

Eukaryot Cell. 2006 Jul;5(7):997-1006. doi: 10.1128/EC.00092-06.

Abstract

The sequencing of the genome of Schizosaccharomyces pombe revealed the presence of a number of genes encoding tandem proteins, some of which are mitochondrial components. One of these proteins (pre-Rsm22-Cox11) consists of a fusion of Rsm22, a component of the mitochondrial ribosome, and Cox11, a factor required for copper insertion into cytochrome oxidase. Since in Saccharomyces cerevisiae, Cox11 is physically attached to the mitochondrial ribosome, it was suggested that the tandem organization of Rsm22-Cox11 is used to covalently tie the mitochondrial ribosome to Cox11 in S. pombe. We report here that pre-Rsm22-Cox11 is matured in two subsequent processing events. First, the mitochondrial presequence is removed. At a later stage of the import process, the Rsm22 and Cox11 domains are separated by cleavage of the mitochondrial processing peptidase at an internal processing site. In vivo data obtained using a tagged version of pre-Rsm22-Cox11 confirmed the proteolytic separation of Cox11 from the Rsm22 domain. Hence, the tandem organization of pre-Rsm22-Cox11 does not give rise to a persistent fusion protein but rather might be used to increase the import efficiency of Cox11 and/or to coordinate expression levels of Rsm22 and Cox11 in S. pombe.

Publication types

  • Comparative Study
  • Research Support, Non-U.S. Gov't

MeSH terms

  • Alternative Splicing
  • Gene Rearrangement
  • Membrane Proteins / metabolism*
  • Mitochondria / metabolism
  • Mitochondrial Proteins / metabolism*
  • Models, Biological
  • Protein Modification, Translational*
  • Protein Sorting Signals
  • Protein Transport
  • Recombinant Fusion Proteins / metabolism
  • Ribosomal Proteins / metabolism*
  • Saccharomyces cerevisiae Proteins / metabolism*
  • Schizosaccharomyces / metabolism*

Substances

  • COX11 protein, S cerevisiae
  • Membrane Proteins
  • Mitochondrial Proteins
  • Protein Sorting Signals
  • Recombinant Fusion Proteins
  • Ribosomal Proteins
  • Rsm22 protein, S cerevisiae
  • Saccharomyces cerevisiae Proteins