Abstract
Human papillomaviruses (HPVs) are a causative factor in over 90% of cervical and 25% of head and neck squamous cell carcinomas (HNSCCs). The C terminus of the high-risk HPV 16 E6 oncoprotein physically associates with and degrades a non-receptor protein tyrosine phosphatase (PTPN13), and PTPN13 loss synergizes with H-Ras(V12) or ErbB2 for invasive growth in vivo. Oral keratinocytes that have lost PTPN13 and express H-Ras(V12) or ErbB2 show enhanced Ras/RAF/MEK/Erk signaling. In co-transfection studies, wild-type PTPN13 inhibited Ras/RAF/MEK/Erk signaling in HEK 293 cells that overexpress ErbB2, EGFR or H-Ras(V12), whereas an enzymatically inactive PTPN13 did not. Twenty percent of HPV-negative HNSCCs had PTPN13 phosphatase mutations that did not inhibit Ras/RAF/MEK/Erk signaling. Inhibition of Ras/RAF/MEK/Erk signaling using MEK inhibitor U0126 blocked anchorage-independent growth in cells lacking PTPN13. These findings show that PTPN13 phosphatase activity has a physiologically significant role in regulating MAP kinase signaling.
Publication types
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Research Support, N.I.H., Extramural
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Research Support, Non-U.S. Gov't
MeSH terms
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Animals
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Butadienes / pharmacology
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Carcinoma, Squamous Cell / enzymology*
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Carcinoma, Squamous Cell / genetics
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Carcinoma, Squamous Cell / pathology
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Carcinoma, Squamous Cell / virology
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Cell Growth Processes / physiology
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Cell Line, Tumor
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Extracellular Signal-Regulated MAP Kinases / metabolism
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Human papillomavirus 16
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Humans
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MAP Kinase Kinase Kinases / antagonists & inhibitors
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MAP Kinase Kinase Kinases / metabolism
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MAP Kinase Signaling System*
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Mice
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Mice, Inbred C57BL
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Nitriles / pharmacology
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Oncogene Proteins, Viral
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Papillomavirus Infections / enzymology*
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Papillomavirus Infections / pathology
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Phosphorylation
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Protein Tyrosine Phosphatase, Non-Receptor Type 13 / deficiency
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Protein Tyrosine Phosphatase, Non-Receptor Type 13 / genetics*
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Protein Tyrosine Phosphatase, Non-Receptor Type 13 / metabolism*
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Receptor, ErbB-2 / genetics*
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Receptor, ErbB-2 / metabolism
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Repressor Proteins
Substances
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Butadienes
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E6 protein, Human papillomavirus type 16
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Nitriles
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Oncogene Proteins, Viral
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Repressor Proteins
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U 0126
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Receptor, ErbB-2
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Extracellular Signal-Regulated MAP Kinases
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MAP Kinase Kinase Kinases
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Protein Tyrosine Phosphatase, Non-Receptor Type 13