The presence of high-molecular-weight viral RNAs interferes with the detection of viral small RNAs

RNA. 2010 May;16(5):1062-7. doi: 10.1261/rna.2049510. Epub 2010 Mar 26.

Abstract

Viral small interfering RNA (siRNA) accumulation in plants is reported to exhibit a strong strand polarity bias, with plus (+) strand siRNAs dominating over minus (-) strand populations. This is of particular interest, as siRNAs processed from double-stranded RNA would be expected to accumulate equivalent amounts of both species. Here, we show that, as reported, (-) strand viral siRNAs are detected at much lower levels than (+) strand-derived species using standard Northern hybridization approaches. However, when total RNA is spiked with in vitro-transcribed antisense viral genomic RNA, (-) strand viral siRNAs are detected at increased levels equivalent to those of (+) strand siRNA. Our results suggest that (+) and (-) strand viral siRNAs accumulate to equivalent levels; however, a proportion of the (-) strand siRNAs are sequestered from the total detectable small RNA population during gel electrophoresis by hybridizing to the high-molecular-weight sense strand viral genomic RNA. Our findings provide a plausible explanation for the observed strand bias of viral siRNA accumulation, and could have wider implications in the analysis of both viral and nonviral small RNA accumulation.

MeSH terms

  • Base Sequence
  • Cucumber Mosaic Virus Satellite / genetics
  • Cucumber Mosaic Virus Satellite / isolation & purification*
  • Cucumber Mosaic Virus Satellite / metabolism
  • Cucumovirus / genetics
  • Cucumovirus / isolation & purification*
  • Cucumovirus / metabolism
  • Electrophoresis, Polyacrylamide Gel
  • Molecular Sequence Data
  • Molecular Weight
  • Nicotiana / virology*
  • RNA Interference
  • RNA Processing, Post-Transcriptional
  • RNA, Small Interfering / genetics
  • RNA, Small Interfering / isolation & purification
  • RNA, Small Interfering / metabolism
  • RNA, Viral / chemistry
  • RNA, Viral / genetics
  • RNA, Viral / isolation & purification*
  • RNA, Viral / metabolism

Substances

  • Cucumber Mosaic Virus Satellite
  • RNA, Small Interfering
  • RNA, Viral