Twin study indicates loss of interaction between microbiota and mucosa of patients with ulcerative colitis

Gastroenterology. 2011 Jul;141(1):227-36. doi: 10.1053/j.gastro.2011.04.011. Epub 2011 Apr 16.

Abstract

Background & aims: Interactions between genetic and environmental factors are believed to be involved in onset and initiation of inflammatory bowel disease. We analyzed the interaction between gastrointestinal mucosal microbiota and host genes in twin pairs discordant for ulcerative colitis (UC) to study the functional interaction between microbiota and mucosal epithelium.

Methods: Biopsy were collected from sigmoid colon of UC patients and their healthy twins (discordant twin pairs) and from twins without UC. Microbiota profiles were determined from analysis of 16S ribosomal DNA libraries; messenger RNA profiles were determined by microarray analysis.

Results: Patients with UC had dysbiotic microbiota, characterized by less bacterial diversity and more Actinobacteria and Proteobacteria than that of their healthy siblings; healthy siblings from discordant twins had more bacteria from the Lachnospiraceae and Ruminococcaceae families than twins who were both healthy. In twins who were both healthy, 34 mucosal transcripts correlated with bacterial genera, whereas only 25 and 11 correlated with bacteria genera in healthy individuals and their twins with UC, respectively. Transcripts related to oxidative and immune responses were differentially expressed between patients with UC and their healthy twins.

Conclusions: The transcriptional profile of the mucosa appears to interact with the colonic microbiota; this interaction appears to be lost in colon of patients with UC. Bacterial functions, such as butyrate production, might affect mucosal gene expression. Patients with UC had different gene expression profiles and lower levels of biodiversity than their healthy twins, as well as unusual aerobic bacteria. Patients with UC had lower percentages of potentially protective bacterial species than their healthy twins.

Publication types

  • Multicenter Study
  • Research Support, Non-U.S. Gov't
  • Twin Study

MeSH terms

  • Adolescent
  • Adult
  • Bacteria / classification
  • Bacteria / genetics
  • Bacteria / growth & development*
  • Biopsy
  • Case-Control Studies
  • Cluster Analysis
  • Colitis, Ulcerative / genetics*
  • Colitis, Ulcerative / microbiology*
  • Colon, Sigmoid / microbiology*
  • DNA, Bacterial / isolation & purification
  • Female
  • Gene Expression Profiling
  • Gene Expression Regulation
  • Germany
  • Heredity
  • Host-Pathogen Interactions / genetics*
  • Humans
  • Intestinal Mucosa / microbiology*
  • Lithuania
  • Male
  • Middle Aged
  • Molecular Sequence Data
  • Principal Component Analysis
  • RNA, Ribosomal, 16S / genetics
  • Ribotyping
  • Twins, Dizygotic / genetics*
  • Twins, Monozygotic / genetics*
  • Young Adult

Substances

  • DNA, Bacterial
  • RNA, Ribosomal, 16S

Associated data

  • GENBANK/HM805116
  • GENBANK/HM805117
  • GENBANK/HM805118
  • GENBANK/HM805119
  • GENBANK/HM805120
  • GENBANK/HM805121
  • GENBANK/HM805122
  • GENBANK/HM805123
  • GENBANK/HM805124
  • GENBANK/HM805125
  • GENBANK/HM805126
  • GENBANK/HM805127
  • GENBANK/HM805128
  • GENBANK/HM805129
  • GENBANK/HM805130
  • GENBANK/HM805131
  • GENBANK/HM805132
  • GENBANK/HM805133
  • GENBANK/HM805134
  • GENBANK/HM805135
  • GENBANK/HM805136
  • GENBANK/HM805137
  • GENBANK/HM805138
  • GENBANK/HM805139
  • GENBANK/HM805140
  • GENBANK/HM805141
  • GENBANK/HM805142
  • GENBANK/HM805143
  • GENBANK/HM805144
  • GENBANK/HM805145
  • GENBANK/HM805146
  • GENBANK/HM805147
  • GENBANK/HM805148
  • GENBANK/HM805149
  • GENBANK/HM805150
  • GENBANK/HM805151
  • GENBANK/HM805152
  • GENBANK/HM805153
  • GENBANK/HM805154
  • GENBANK/HM805155
  • GENBANK/HM805156
  • GENBANK/HM805157
  • GENBANK/HM805158
  • GENBANK/HM805159
  • GENBANK/HM805160
  • GENBANK/HM805161
  • GENBANK/HM805162
  • GENBANK/HM805163
  • GENBANK/HM805164
  • GENBANK/HM805165
  • GENBANK/HM805166
  • GENBANK/HM805167
  • GENBANK/HM805168
  • GENBANK/HM805169
  • GENBANK/HM805170
  • GENBANK/HM805171
  • GENBANK/HM805172
  • GENBANK/HM805173
  • GENBANK/HM805174
  • GENBANK/HM805175
  • GENBANK/HM805176
  • GENBANK/HM805177
  • GENBANK/HM805178
  • GENBANK/HM805179
  • GENBANK/HM805180
  • GENBANK/HM805181
  • GENBANK/HM805182
  • GENBANK/HM805183
  • GENBANK/HM805184
  • GENBANK/HM805185
  • GENBANK/HM805186
  • GENBANK/HM805187
  • GENBANK/HM805188
  • GENBANK/HM805189
  • GENBANK/HM805190
  • GENBANK/HM805191
  • GENBANK/HM805192
  • GENBANK/HM805193
  • GENBANK/HM805194
  • GENBANK/HM805195
  • GENBANK/HM805196
  • GENBANK/HM805197
  • GENBANK/HM805198
  • GENBANK/HM805199
  • GENBANK/HM805200
  • GENBANK/HM805201
  • GENBANK/HM805202
  • GENBANK/HM805203
  • GENBANK/HM805204
  • GENBANK/HM805205
  • GENBANK/HM805206
  • GENBANK/HM805207
  • GENBANK/HM805208
  • GENBANK/HM805209
  • GENBANK/HM805210
  • GENBANK/HM805211
  • GENBANK/HM805212
  • GENBANK/HM805213
  • GENBANK/HM805214
  • GENBANK/HM805215
  • GENBANK/HM805216
  • GENBANK/HM805217
  • GENBANK/HM805218
  • GENBANK/HM805219
  • GENBANK/HM805220
  • GENBANK/HM805221
  • GENBANK/HM805222
  • GENBANK/HM805223
  • GENBANK/HM805224
  • GENBANK/HM805225
  • GENBANK/HM805226
  • GENBANK/HM805227
  • GENBANK/HM805228
  • GENBANK/HM805229
  • GENBANK/HM805230
  • GENBANK/HM805231
  • GENBANK/HM805232
  • GENBANK/HM805233
  • GENBANK/HM805234
  • GENBANK/HM805235
  • GENBANK/HM805236
  • GENBANK/HM805237
  • GENBANK/HM805238
  • GENBANK/HM805239
  • GENBANK/HM805240
  • GENBANK/HM805241
  • GENBANK/HM805242
  • GENBANK/HM805243
  • GENBANK/HM805244
  • GENBANK/HM805245
  • GENBANK/HM805246
  • GENBANK/HM805247
  • GENBANK/HM805248
  • GENBANK/HM805249
  • GENBANK/HM805250
  • GENBANK/HM805251
  • GENBANK/HM805252
  • GENBANK/HM805253
  • GENBANK/HM805254
  • GENBANK/HM805255
  • GENBANK/HM805256
  • GENBANK/HM805257
  • GENBANK/HM805258
  • GENBANK/HM805259
  • GENBANK/HM805260
  • GENBANK/HM805261
  • GENBANK/HM805262
  • GENBANK/HM805263
  • GENBANK/HM805264
  • GENBANK/HM805265
  • GENBANK/HM805266
  • GENBANK/HM805267
  • GENBANK/HM805268
  • GENBANK/HM805269
  • GENBANK/HM805270
  • GENBANK/HM805271
  • GENBANK/HM805272
  • GENBANK/HM805273
  • GENBANK/HM805274
  • GENBANK/HM805275
  • GENBANK/HM805276
  • GENBANK/HM805277
  • GENBANK/HM805278
  • GENBANK/HM805279
  • GENBANK/HM805280
  • GENBANK/HM805281
  • GENBANK/HM805282
  • GENBANK/HM805283
  • GENBANK/HM805284
  • GENBANK/HM805285
  • GENBANK/HM805286
  • GENBANK/HM805287
  • GENBANK/HM805288
  • GENBANK/HM805289
  • GENBANK/HM805290
  • GENBANK/HM805291
  • GENBANK/HM805292
  • GENBANK/HM805293
  • GENBANK/HM805294
  • GENBANK/HM805295
  • GENBANK/HM805296
  • GENBANK/HM805297
  • GENBANK/HM805298
  • GENBANK/HM805299
  • GENBANK/HM805300
  • GENBANK/HM805301
  • GENBANK/HM805302
  • GENBANK/HM805303
  • GENBANK/HM805304
  • GENBANK/HM805305
  • GENBANK/HM805306
  • GENBANK/HM805307
  • GENBANK/HM805308
  • GENBANK/HM805309
  • GENBANK/HM805310
  • GENBANK/HM805311
  • GENBANK/HM805312
  • GENBANK/HM805313
  • GENBANK/HM805314
  • GENBANK/HM805315
  • GENBANK/HM805316
  • GENBANK/HM805317
  • GENBANK/HM805318
  • GENBANK/HM805319
  • GENBANK/HM805320
  • GENBANK/HM805321
  • GENBANK/HM805322
  • GENBANK/HM805323
  • GENBANK/HM805324
  • GENBANK/HM805325
  • GENBANK/HM805326
  • GENBANK/HM805327
  • GENBANK/HM805328
  • GENBANK/HM805329
  • GENBANK/HM805330
  • GENBANK/HM805331
  • GENBANK/HM805332
  • GENBANK/HM805333
  • GENBANK/HM805334
  • GENBANK/HM805335
  • GENBANK/HM805336
  • GENBANK/HM805337
  • GENBANK/HM805338
  • GENBANK/HM805339
  • GENBANK/HM805340
  • GENBANK/HM805341
  • GENBANK/HM805342
  • GENBANK/HM805343
  • GENBANK/HM805344
  • GENBANK/HM805345
  • GENBANK/HM805346
  • GENBANK/HM805347
  • GENBANK/HM805348
  • GENBANK/HM805349
  • GENBANK/HM805350
  • GENBANK/HM805351
  • GENBANK/HM805352
  • GENBANK/HM805353
  • GENBANK/HM805354
  • GENBANK/HM805355
  • GENBANK/HM805356
  • GENBANK/HM805357
  • GENBANK/HM805358
  • GENBANK/HM805359
  • GENBANK/HM805360
  • GENBANK/HM805361
  • GENBANK/HM805362
  • GENBANK/HM805363
  • GENBANK/HM805364
  • GENBANK/HM805365
  • GENBANK/HM805366
  • GENBANK/HM805367
  • GENBANK/HM805368
  • GENBANK/HM805369
  • GENBANK/HM805370
  • GENBANK/HM805371
  • GENBANK/HM805372
  • GENBANK/HM805373
  • GENBANK/HM805374
  • GENBANK/HM805375
  • GENBANK/HM805376
  • GENBANK/HM805377
  • GENBANK/HM805378
  • GENBANK/HM805379
  • GENBANK/HM805380
  • GENBANK/HM805381
  • GENBANK/HM805382
  • GENBANK/HM805383
  • GENBANK/HM805384
  • GENBANK/HM805385
  • GENBANK/HM805386
  • GENBANK/HM805387
  • GENBANK/HM805388
  • GENBANK/HM805389
  • GENBANK/HM805390
  • GENBANK/HM805391
  • GENBANK/HM805392
  • GENBANK/HM805393
  • GENBANK/HM805394
  • GENBANK/HM805395
  • GENBANK/HM805396
  • GENBANK/HM805397
  • GENBANK/HM805398
  • GENBANK/HM805399
  • GENBANK/HM805400
  • GENBANK/HM805401
  • GENBANK/HM805402
  • GENBANK/HM805403
  • GENBANK/HM805404
  • GENBANK/HM805405
  • GENBANK/HM805406
  • GENBANK/HM805407
  • GENBANK/HM805408
  • GENBANK/HM805409
  • GENBANK/HM805410
  • GENBANK/HM805411
  • GENBANK/HM805412
  • GENBANK/HM805413
  • GENBANK/HM805414
  • GENBANK/HM805415
  • GENBANK/HM805416
  • GENBANK/HM805417
  • GENBANK/HM805418
  • GENBANK/HM805419
  • GENBANK/HM805420
  • GENBANK/HM805421
  • GENBANK/HM805422
  • GENBANK/HM805423
  • GENBANK/HM805424
  • GENBANK/HM805425
  • GENBANK/HM805426
  • GENBANK/HM805427
  • GENBANK/HM805428
  • GENBANK/HM805429
  • GENBANK/HM805430
  • GENBANK/HM805431
  • GENBANK/HM805432
  • GENBANK/HM805433
  • GENBANK/HM805434
  • GENBANK/HM805435
  • GENBANK/HM805436
  • GENBANK/HM805437
  • GENBANK/HM805438
  • GENBANK/HM805439
  • GENBANK/HM805440
  • GENBANK/HM805441
  • GENBANK/HM805442
  • GENBANK/HM805443
  • GENBANK/HM805444
  • GENBANK/HM805445
  • GENBANK/HM805446
  • GENBANK/HM805447
  • GENBANK/HM805448
  • GENBANK/HM805449
  • GENBANK/HM805450
  • GENBANK/HM805451
  • GENBANK/HM805452
  • GENBANK/HM805453
  • GENBANK/HM805454
  • GENBANK/HM805455
  • GENBANK/HM805456
  • GENBANK/HM805457
  • GENBANK/HM805458
  • GENBANK/HM805459
  • GENBANK/HM805460
  • GENBANK/HM805461
  • GENBANK/HM805462
  • GENBANK/HM805463
  • GENBANK/HM805464
  • GENBANK/HM805465
  • GENBANK/HM805466
  • GENBANK/HM805467
  • GENBANK/HM805468
  • GENBANK/HM805469
  • GENBANK/HM805470
  • GENBANK/HM805471
  • GENBANK/HM805472
  • GENBANK/HM805473
  • GENBANK/HM805474
  • GENBANK/HM805475
  • GENBANK/HM805476
  • GENBANK/HM805477
  • GENBANK/HM805478
  • GENBANK/HM805479
  • GENBANK/HM805480
  • GENBANK/HM805481
  • GENBANK/HM805482
  • GENBANK/HM805483
  • GENBANK/HM805484
  • GENBANK/HM805485
  • GENBANK/HM805486
  • GENBANK/HM805487
  • GENBANK/HM805488
  • GENBANK/HM805489
  • GENBANK/HM805490
  • GENBANK/HM805491
  • GENBANK/HM805492
  • GENBANK/HM805493
  • GENBANK/HM805494
  • GENBANK/HM805495
  • GENBANK/HM805496
  • GENBANK/HM805497
  • GENBANK/HM805498
  • GENBANK/HM805499
  • GENBANK/HM805500
  • GENBANK/HM805501
  • GENBANK/HM805502
  • GENBANK/HM805503
  • GENBANK/HM805504
  • GENBANK/HM805505
  • GENBANK/HM805506
  • GENBANK/HM805507
  • GENBANK/HM805508
  • GENBANK/HM805509
  • GENBANK/HM805510
  • GENBANK/HM805511
  • GENBANK/HM805512
  • GENBANK/HM805513
  • GENBANK/HM805514
  • GENBANK/HM805515
  • GENBANK/HM805516
  • GENBANK/HM805517
  • GENBANK/HM805518
  • GENBANK/HM805519
  • GENBANK/HM805520
  • GENBANK/HM805521
  • GENBANK/HM805522
  • GENBANK/HM805523
  • GENBANK/HM805524
  • GENBANK/HM805525
  • GENBANK/HM805526
  • GENBANK/HM805527
  • GENBANK/HM805528
  • GENBANK/HM805529
  • GENBANK/HM805530
  • GENBANK/HM805531
  • GENBANK/HM805532
  • GENBANK/HM805533
  • GENBANK/HM805534
  • GENBANK/HM805535
  • GENBANK/HM805536
  • GENBANK/HM805537
  • GENBANK/HM805538
  • GENBANK/HM805539
  • GENBANK/HM805540
  • GENBANK/HM805541
  • GENBANK/HM805542
  • GENBANK/HM805543
  • GENBANK/HM805544
  • GENBANK/HM805545
  • GENBANK/HM805546
  • GENBANK/HM805547
  • GENBANK/HM805548
  • GENBANK/HM805549
  • GENBANK/HM805550
  • GENBANK/HM805551
  • GENBANK/HM805552
  • GENBANK/HM805553
  • GENBANK/HM805554
  • GENBANK/HM805555
  • GENBANK/HM805556
  • GENBANK/HM805557
  • GENBANK/HM805558
  • GENBANK/HM805559
  • GENBANK/HM805560
  • GENBANK/HM805561
  • GENBANK/HM805562
  • GENBANK/HM805563
  • GENBANK/HM805564
  • GENBANK/HM805565
  • GENBANK/HM805566
  • GENBANK/HM805567
  • GENBANK/HM805568
  • GENBANK/HM805569
  • GENBANK/HM805570
  • GENBANK/HM805571
  • GENBANK/HM805572
  • GENBANK/HM805573
  • GENBANK/HM805574
  • GENBANK/HM805575
  • GENBANK/HM805576
  • GENBANK/HM805577
  • GENBANK/HM805578
  • GENBANK/HM805579
  • GENBANK/HM805580
  • GENBANK/HM805581
  • GENBANK/HM805582
  • GENBANK/HM805583
  • GENBANK/HM805584
  • GENBANK/HM805585
  • GENBANK/HM805586
  • GENBANK/HM805587
  • GENBANK/HM805588
  • GENBANK/HM805589
  • GENBANK/HM805590
  • GENBANK/HM805591
  • GENBANK/HM805592
  • GENBANK/HM805593
  • GENBANK/HM805594
  • GENBANK/HM805595
  • GENBANK/HM805596
  • GENBANK/HM805597
  • GENBANK/HM805598
  • GENBANK/HM805599
  • GENBANK/HM805600
  • GENBANK/HM805601
  • GENBANK/HM805602
  • GENBANK/HM805603
  • GENBANK/HM805604
  • GENBANK/HM805605
  • GENBANK/HM805606
  • GENBANK/HM805607
  • GENBANK/HM805608
  • GENBANK/HM805609
  • GENBANK/HM805610
  • GENBANK/HM805611
  • GENBANK/HM805612
  • GENBANK/HM805613
  • GENBANK/HM805614
  • GENBANK/HM805615
  • GENBANK/HM805616
  • GENBANK/HM805617
  • GENBANK/HM805618
  • GENBANK/HM805619
  • GENBANK/HM805620
  • GENBANK/HM805621
  • GENBANK/HM805622
  • GENBANK/HM805623
  • GENBANK/HM805624
  • GENBANK/HM805625
  • GENBANK/HM805626
  • GENBANK/HM805627
  • GENBANK/HM805628
  • GENBANK/HM805629
  • GENBANK/HM805630
  • GENBANK/HM805631
  • GENBANK/HM805632
  • GENBANK/HM805633
  • GENBANK/HM805634
  • GENBANK/HM805635
  • GENBANK/HM805636
  • GENBANK/HM805637
  • GENBANK/HM805638
  • GENBANK/HM805639
  • GENBANK/HM805640
  • GENBANK/HM805641
  • GENBANK/HM805642
  • GENBANK/HM805643
  • GENBANK/HM805644
  • GENBANK/HM805645
  • GENBANK/HM805646
  • GENBANK/HM805647
  • GENBANK/HM805648
  • GENBANK/HM805649
  • GENBANK/HM805650
  • GENBANK/HM805651
  • GENBANK/HM805652
  • GENBANK/HM805653
  • GENBANK/HM805654
  • GENBANK/HM805655
  • GENBANK/HM805656
  • GENBANK/HM805657
  • GENBANK/HM805658
  • GENBANK/HM805659
  • GENBANK/HM805660
  • GENBANK/HM805661
  • GENBANK/HM805662
  • GENBANK/HM805663
  • GENBANK/HM805664
  • GENBANK/HM805665
  • GENBANK/HM805666
  • GENBANK/HM805667
  • GENBANK/HM805668
  • GENBANK/HM805669
  • GENBANK/HM805670
  • GENBANK/HM805671
  • GENBANK/HM805672
  • GENBANK/HM805673
  • GENBANK/HM805674
  • GENBANK/HM805675
  • GENBANK/HM805676
  • GENBANK/HM805677
  • GENBANK/HM805678
  • GENBANK/HM805679
  • GENBANK/HM805680
  • GENBANK/HM805681
  • GENBANK/HM805682
  • GENBANK/HM805683
  • GENBANK/HM805684
  • GENBANK/HM805685
  • GENBANK/HM805686
  • GENBANK/HM805687
  • GENBANK/HM805688
  • GENBANK/HM805689
  • GENBANK/HM805690
  • GENBANK/HM805691
  • GENBANK/HM805692
  • GENBANK/HM805693
  • GENBANK/HM805694
  • GENBANK/HM805695
  • GENBANK/HM805696
  • GENBANK/HM805697
  • GENBANK/HM805698
  • GENBANK/HM805699
  • GENBANK/HM805700
  • GENBANK/HM805701
  • GENBANK/HM805702
  • GENBANK/HM805703
  • GENBANK/HM805704
  • GENBANK/HM805705
  • GENBANK/HM805706
  • GENBANK/HM805707
  • GENBANK/HM805708
  • GENBANK/HM805709
  • GENBANK/HM805710
  • GENBANK/HM805711
  • GENBANK/HM805712
  • GENBANK/HM805713
  • GENBANK/HM805714
  • GENBANK/HM805715
  • GENBANK/HM805716
  • GENBANK/HM805717
  • GENBANK/HM805718
  • GENBANK/HM805719
  • GENBANK/HM805720
  • GENBANK/HM805721
  • GENBANK/HM805722
  • GENBANK/HM805723
  • GENBANK/HM805724
  • GENBANK/HM805725
  • GENBANK/HM805726
  • GENBANK/HM805727
  • GENBANK/HM805728
  • GENBANK/HM805729
  • GENBANK/HM805730
  • GENBANK/HM805731
  • GENBANK/HM805732
  • GENBANK/HM805733
  • GENBANK/HM805734
  • GENBANK/HM805735
  • GENBANK/HM805736
  • GENBANK/HM805737
  • GENBANK/HM805738
  • GENBANK/HM805739
  • GENBANK/HM805740
  • GENBANK/HM805741
  • GENBANK/HM805742
  • GENBANK/HM805743
  • GENBANK/HM805744
  • GENBANK/HM805745
  • GENBANK/HM805746
  • GENBANK/HM805747
  • GENBANK/HM805748
  • GENBANK/HM805749
  • GENBANK/HM805750
  • GENBANK/HM805751
  • GENBANK/HM805752
  • GENBANK/HM805753
  • GENBANK/HM805754
  • GENBANK/HM805755
  • GENBANK/HM805756
  • GENBANK/HM805757
  • GENBANK/HM805758
  • GENBANK/HM805759
  • GENBANK/HM805760
  • GENBANK/HM805761
  • GENBANK/HM805762
  • GENBANK/HM805763
  • GENBANK/HM805764
  • GENBANK/HM805765
  • GENBANK/HM805766
  • GENBANK/HM805767
  • GENBANK/HM805768
  • GENBANK/HM805769
  • GENBANK/HM805770
  • GENBANK/HM805771
  • GENBANK/HM805772
  • GENBANK/HM805773
  • GENBANK/HM805774
  • GENBANK/HM805775
  • GENBANK/HM805776
  • GENBANK/HM805777
  • GENBANK/HM805778
  • GENBANK/HM805779
  • GENBANK/HM805780
  • GENBANK/HM805781
  • GENBANK/HM805782
  • GENBANK/HM805783
  • GENBANK/HM805784
  • GENBANK/HM805785
  • GENBANK/HM805786
  • GENBANK/HM805787
  • GENBANK/HM805788
  • GENBANK/HM805789
  • GENBANK/HM805790
  • GENBANK/HM805791
  • GENBANK/HM805792
  • GENBANK/HM805793
  • GENBANK/HM805794
  • GENBANK/HM805795
  • GENBANK/HM805796
  • GENBANK/HM805797
  • GENBANK/HM805798
  • GENBANK/HM805799
  • GENBANK/HM805800
  • GENBANK/HM805801
  • GENBANK/HM805802
  • GENBANK/HM805803
  • GENBANK/HM805804
  • GENBANK/HM805805
  • GENBANK/HM805806
  • GENBANK/HM805807
  • GENBANK/HM805808
  • GENBANK/HM805809
  • GENBANK/HM805810
  • GENBANK/HM805811
  • GENBANK/HM805812
  • GENBANK/HM805813
  • GENBANK/HM805814
  • GENBANK/HM805815
  • GENBANK/HM805816
  • GENBANK/HM805817
  • GENBANK/HM805818
  • GENBANK/HM805819
  • GENBANK/HM805820
  • GENBANK/HM805821
  • GENBANK/HM805822
  • GENBANK/HM805823
  • GENBANK/HM805824
  • GENBANK/HM805825
  • GENBANK/HM805826
  • GENBANK/HM805827
  • GENBANK/HM805828
  • GENBANK/HM805829
  • GENBANK/HM805830
  • GENBANK/HM805831
  • GENBANK/HM805832
  • GENBANK/HM805833
  • GENBANK/HM805834
  • GENBANK/HM805835
  • GENBANK/HM805836
  • GENBANK/HM805837
  • GENBANK/HM805838
  • GENBANK/HM805839
  • GENBANK/HM805840
  • GENBANK/HM805841
  • GENBANK/HM805842
  • GENBANK/HM805843
  • GENBANK/HM805844
  • GENBANK/HM805845
  • GENBANK/HM805846
  • GENBANK/HM805847
  • GENBANK/HM805848
  • GENBANK/HM805849
  • GENBANK/HM805850
  • GENBANK/HM805851
  • GENBANK/HM805852
  • GENBANK/HM805853
  • GENBANK/HM805854
  • GENBANK/HM805855
  • GENBANK/HM805856
  • GENBANK/HM805857
  • GENBANK/HM805858
  • GENBANK/HM805859
  • GENBANK/HM805860
  • GENBANK/HM805861
  • GENBANK/HM805862
  • GENBANK/HM805863
  • GENBANK/HM805864
  • GENBANK/HM805865
  • GENBANK/HM805866
  • GENBANK/HM805867
  • GENBANK/HM805868
  • GENBANK/HM805869
  • GENBANK/HM805870
  • GENBANK/HM805871
  • GENBANK/HM805872
  • GENBANK/HM805873
  • GENBANK/HM805874
  • GENBANK/HM805875
  • GENBANK/HM805876
  • GENBANK/HM805877
  • GENBANK/HM805878
  • GENBANK/HM805879
  • GENBANK/HM805880
  • GENBANK/HM805881
  • GENBANK/HM805882
  • GENBANK/HM805883
  • GENBANK/HM805884
  • GENBANK/HM805885
  • GENBANK/HM805886
  • GENBANK/HM805887
  • GENBANK/HM805888
  • GENBANK/HM805889
  • GENBANK/HM805890
  • GENBANK/HM805891
  • GENBANK/HM805892
  • GENBANK/HM805893
  • GENBANK/HM805894
  • GENBANK/HM805895
  • GENBANK/HM805896
  • GENBANK/HM805897
  • GENBANK/HM805898
  • GENBANK/HM805899
  • GENBANK/HM805900
  • GENBANK/HM805901
  • GENBANK/HM805902
  • GENBANK/HM805903
  • GENBANK/HM805904
  • GENBANK/HM805905
  • GENBANK/HM805906
  • GENBANK/HM805907
  • GENBANK/HM805908
  • GENBANK/HM805909
  • GENBANK/HM805910
  • GENBANK/HM805911
  • GENBANK/HM805912
  • GENBANK/HM805913
  • GENBANK/HM805914
  • GENBANK/HM805915
  • GENBANK/HM805916
  • GENBANK/HM805917
  • GENBANK/HM805918
  • GENBANK/HM805919
  • GENBANK/HM805920
  • GENBANK/HM805921
  • GENBANK/HM805922
  • GENBANK/HM805923
  • GENBANK/HM805924
  • GENBANK/HM805925
  • GENBANK/HM805926
  • GENBANK/HM805927
  • GENBANK/HM805928
  • GENBANK/HM805929
  • GENBANK/HM805930
  • GENBANK/HM805931
  • GENBANK/HM805932
  • GENBANK/HM805933
  • GENBANK/HM805934
  • GENBANK/HM805935
  • GENBANK/HM805936
  • GENBANK/HM805937
  • GENBANK/HM805938
  • GENBANK/HM805939
  • GENBANK/HM805940
  • GENBANK/HM805941
  • GENBANK/HM805942
  • GENBANK/HM805943
  • GENBANK/HM805944
  • GENBANK/HM805945
  • GENBANK/HM805946
  • GENBANK/HM805947
  • GENBANK/HM805948
  • GENBANK/HM805949
  • GENBANK/HM805950
  • GENBANK/HM805951
  • GENBANK/HM805952
  • GENBANK/HM805953
  • GENBANK/HM805954
  • GENBANK/HM805955
  • GENBANK/HM805956
  • GENBANK/HM805957
  • GENBANK/HM805958
  • GENBANK/HM805959
  • GENBANK/HM805960
  • GENBANK/HM805961
  • GENBANK/HM805962
  • GENBANK/HM805963
  • GENBANK/HM805964
  • GENBANK/HM805965
  • GENBANK/HM805966
  • GENBANK/HM805967
  • GENBANK/HM805968
  • GENBANK/HM805969
  • GENBANK/HM805970
  • GENBANK/HM805971
  • GENBANK/HM805972
  • GENBANK/HM805973
  • GENBANK/HM805974
  • GENBANK/HM805975
  • GENBANK/HM805976
  • GENBANK/HM805977
  • GENBANK/HM805978
  • GENBANK/HM805979
  • GENBANK/HM805980
  • GENBANK/HM805981
  • GENBANK/HM805982
  • GENBANK/HM805983
  • GENBANK/HM805984
  • GENBANK/HM805985
  • GENBANK/HM805986
  • GENBANK/HM805987
  • GENBANK/HM805988
  • GENBANK/HM805989
  • GENBANK/HM805990
  • GENBANK/HM805991
  • GENBANK/HM805992
  • GENBANK/HM805993
  • GENBANK/HM805994
  • GENBANK/HM805995
  • GENBANK/HM805996
  • GENBANK/HM805997
  • GENBANK/HM805998
  • GENBANK/HM805999
  • GENBANK/HM806000
  • GENBANK/HM806001
  • GENBANK/HM806002
  • GENBANK/HM806003
  • GENBANK/HM806004
  • GENBANK/HM806005
  • GENBANK/HM806006
  • GENBANK/HM806007
  • GENBANK/HM806008
  • GENBANK/HM806009
  • GENBANK/HM806010
  • GENBANK/HM806011
  • GENBANK/HM806012
  • GENBANK/HM806013
  • GENBANK/HM806014
  • GENBANK/HM806015
  • GENBANK/HM806016
  • GENBANK/HM806017
  • GENBANK/HM806018
  • GENBANK/HM806019
  • GENBANK/HM806020
  • GENBANK/HM806021
  • GENBANK/HM806022
  • GENBANK/HM806023
  • GENBANK/HM806024
  • GENBANK/HM806025
  • GENBANK/HM806026
  • GENBANK/HM806027
  • GENBANK/HM806028
  • GENBANK/HM806029
  • GENBANK/HM806030
  • GENBANK/HM806031
  • GENBANK/HM806032
  • GENBANK/HM806033
  • GENBANK/HM806034
  • GENBANK/HM806035
  • GENBANK/HM806036
  • GENBANK/HM806037
  • GENBANK/HM806038
  • GENBANK/HM806039
  • GENBANK/HM806040
  • GENBANK/HM806041
  • GENBANK/HM806042
  • GENBANK/HM806043
  • GENBANK/HM806044
  • GENBANK/HM806045
  • GENBANK/HM806046
  • GENBANK/HM806047
  • GENBANK/HM806048
  • GENBANK/HM806049
  • GENBANK/HM806050
  • GENBANK/HM806051
  • GENBANK/HM806052
  • GENBANK/HM806053
  • GENBANK/HM806054
  • GENBANK/HM806055
  • GENBANK/HM806056
  • GENBANK/HM806057
  • GENBANK/HM806058
  • GENBANK/HM806059
  • GENBANK/HM806060
  • GENBANK/HM806061
  • GENBANK/HM806062
  • GENBANK/HM806063
  • GENBANK/HM806064
  • GENBANK/HM806065
  • GENBANK/HM806066
  • GENBANK/HM806067
  • GENBANK/HM806068
  • GENBANK/HM806069
  • GENBANK/HM806070
  • GENBANK/HM806071
  • GENBANK/HM806072
  • GENBANK/HM806073
  • GENBANK/HM806074
  • GENBANK/HM806075
  • GENBANK/HM806076
  • GENBANK/HM806077
  • GENBANK/HM806078
  • GENBANK/HM806079
  • GENBANK/HM806080
  • GENBANK/HM806081
  • GENBANK/HM806082
  • GENBANK/HM806083
  • GENBANK/HM806084
  • GENBANK/HM806085
  • GENBANK/HM806086
  • GENBANK/HM806087
  • GENBANK/HM806088
  • GENBANK/HM806089
  • GENBANK/HM806090
  • GENBANK/HM806091
  • GENBANK/HM806092
  • GENBANK/HM806093
  • GENBANK/HM806094
  • GENBANK/HM806095
  • GENBANK/HM806096
  • GENBANK/HM806097
  • GENBANK/HM806098
  • GENBANK/HM806099
  • GENBANK/HM806100
  • GENBANK/HM806101
  • GENBANK/HM806102
  • GENBANK/HM806103
  • GENBANK/HM806104
  • GENBANK/HM806105
  • GENBANK/HM806106
  • GENBANK/HM806107
  • GENBANK/HM806108
  • GENBANK/HM806109
  • GENBANK/HM806110
  • GENBANK/HM806111
  • GENBANK/HM806112
  • GENBANK/HM806113
  • GEO/GSE22619
  • GEO/GSE7821