Chromosomes without a 30-nm chromatin fiber

Nucleus. 2012 Sep-Oct;3(5):404-10. doi: 10.4161/nucl.21222. Epub 2012 Jul 31.

Abstract

How is a long strand of genomic DNA packaged into a mitotic chromosome or nucleus? The nucleosome fiber (beads-on-a-string), in which DNA is wrapped around core histones, has long been assumed to be folded into a 30-nm chromatin fiber, and a further helically folded larger fiber. However, when frozen hydrated human mitotic cells were observed using cryoelectron microscopy, no higher-order structures that included 30-nm chromatin fibers were found. To investigate the bulk structure of mitotic chromosomes further, we performed small-angle X-ray scattering (SAXS), which can detect periodic structures in noncrystalline materials in solution. The results were striking: no structural feature larger than 11 nm was detected, even at a chromosome-diameter scale (~1 μm). We also found a similar scattering pattern in interphase nuclei of HeLa cells in the range up to ~275 nm. Our findings suggest a common structural feature in interphase and mitotic chromatins: compact and irregular folding of nucleosome fibers occurs without a 30-nm chromatin structure.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Cell Nucleus / metabolism
  • Chromatin / chemistry*
  • Chromatin / metabolism
  • Chromosome Structures / chemistry*
  • Chromosome Structures / metabolism
  • HeLa Cells
  • Histones / chemistry
  • Histones / metabolism
  • Humans
  • Interphase
  • Mitosis
  • Nucleosomes / chemistry
  • Nucleosomes / metabolism
  • Scattering, Small Angle
  • X-Ray Diffraction

Substances

  • Chromatin
  • Histones
  • Nucleosomes