[The properties of four C-terminal carbohydrate-binding modules (CBM4) of laminarinase Lic16A of Clostridium thermocellum]

Mol Biol (Mosk). 2012 Nov-Dec;46(6):915-21.
[Article in Russian]

Abstract

At the C-terminus of multimodular laminarinase Lic16A Clostridium thermocellum four carbohydrate-binding modules (CBM), belonging to family 4, were found. The isolated CBM - CBM4_1, CBM4_2, CBM4_3, CBM4_4 and the tandem CBM4_(1-4) were obtained. None of the recombinant proteins did have the affinity to soluble beta-1,3-1,4-glucans--laminarin and lihenan--the main specific substrates of Licl6A. All modules, except CBM4_4, had the ability to bind bacterial crystalline cellulose, that was atypical for the family 4 CBMs. We found that all CBMs 4 of Licl6A had affinity for xylan, chitin, beta-glucan from yeast cell wall and Avicel, while CBM4_3 and CBM4_4 had additional affinity to chitosan. The tandem CBM4_(1-4) had the highest affinity to yeast cell wall beta-glucan, avicel and pustulan. The binding constants for these substrates were about 100 times higher than that of the individual modules, suggesting a synergy in the process of absorption to these polysaccharides. This finding helps to explain the evolutionary process of CBM multiplication.

Publication types

  • English Abstract
  • Research Support, Non-U.S. Gov't

MeSH terms

  • Bacterial Proteins / chemistry*
  • Bacterial Proteins / genetics
  • Bacterial Proteins / metabolism
  • Cellulases / chemistry*
  • Cellulases / genetics
  • Cellulases / metabolism
  • Clostridium thermocellum / enzymology*
  • Clostridium thermocellum / genetics
  • Evolution, Molecular*
  • Polysaccharides / chemistry*
  • Polysaccharides / genetics
  • Polysaccharides / metabolism
  • Protein Structure, Tertiary
  • Substrate Specificity / physiology

Substances

  • Bacterial Proteins
  • Polysaccharides
  • Cellulases