Heat shock cognate 71 (HSC71) regulates cellular antiviral response by impairing formation of VISA aggregates

Protein Cell. 2013 May;4(5):373-82. doi: 10.1007/s13238-013-3902-3. Epub 2013 Apr 30.

Abstract

In response to viral infection, RIG-I-like RNA helicases detect viral RNA and signal through the mitochondrial adapter protein VISA. VISA activation leads to rapid activation of transcription factors IRF3 and NF-κB, which collaborate to induce transcription of type I interferon (IFN) genes and cellular antiviral response. It has been demonstrated that VISA is activated by forming prion-like aggregates. However, how this process is regulated remains unknown. Here we show that overexpression of HSC71 resulted in potent inhibition of virus-triggered transcription of IFNB1 gene and cellular antiviral response. Consistently, knockdown of HSC71 had opposite effects. HSC71 interacted with VISA, and negatively regulated virus-triggered VISA aggregation. These findings suggest that HSC71 functions as a check against VISA-mediated antiviral response.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Adaptor Proteins, Signal Transducing / biosynthesis*
  • Adaptor Proteins, Signal Transducing / chemistry
  • Adaptor Proteins, Signal Transducing / genetics*
  • Adaptor Proteins, Signal Transducing / metabolism
  • Cell Aggregation / genetics
  • GPI-Linked Proteins / metabolism
  • Gene Knockdown Techniques
  • HEK293 Cells
  • HSC70 Heat-Shock Proteins / genetics*
  • HSC70 Heat-Shock Proteins / metabolism
  • Heat-Shock Response / genetics
  • Humans
  • Interferon Regulatory Factor-3 / genetics
  • Interferon Regulatory Factor-3 / metabolism
  • Interferon-beta / genetics
  • NF-kappa B / genetics
  • Prions / metabolism
  • Receptors, Retinoic Acid / metabolism
  • Viruses / drug effects
  • Viruses / metabolism*
  • Viruses / pathogenicity

Substances

  • Adaptor Proteins, Signal Transducing
  • GPI-Linked Proteins
  • HSC70 Heat-Shock Proteins
  • HSPA8 protein, human
  • IRF3 protein, human
  • Interferon Regulatory Factor-3
  • MAVS protein, human
  • NF-kappa B
  • PLAAT4 protein, human
  • PRND protein, human
  • Prions
  • Receptors, Retinoic Acid
  • Interferon-beta