Modulation of lipid kinase PI4KIIα activity and lipid raft association of presenilin 1 underlies γ-secretase inhibition by ginsenoside (20S)-Rg3

J Biol Chem. 2013 Jul 19;288(29):20868-20882. doi: 10.1074/jbc.M112.445734. Epub 2013 May 30.

Abstract

Amyloid β-peptide (Aβ) pathology is an invariant feature of Alzheimer disease, preceding any detectable clinical symptoms by more than a decade. To this end, we seek to identify agents that can reduce Aβ levels in the brain via novel mechanisms. We found that (20S)-Rg3, a triterpene natural compound known as ginsenoside, reduced Aβ levels in cultured primary neurons and in the brains of a mouse model of Alzheimer disease. The (20S)-Rg3 treatment induced a decrease in the association of presenilin 1 (PS1) fragments with lipid rafts where catalytic components of the γ-secretase complex are enriched. The Aβ-lowering activity of (20S)-Rg3 directly correlated with increased activity of phosphatidylinositol 4-kinase IIα (PI4KIIα), a lipid kinase that mediates the rate-limiting step in phosphatidylinositol 4,5-bisphosphate synthesis. PI4KIIα overexpression recapitulated the effects of (20S)-Rg3, whereas reduced expression of PI4KIIα abolished the Aβ-reducing activity of (20S)-Rg3 in neurons. Our results substantiate an important role for PI4KIIα and phosphoinositide modulation in γ-secretase activity and Aβ biogenesis.

Keywords: Alzheimer Disease; Lipid Raft; Lipids; Natural Products; Phosphatidylinositol; Presenilin.

Publication types

  • Research Support, N.I.H., Extramural
  • Research Support, Non-U.S. Gov't

MeSH terms

  • Amyloid Precursor Protein Secretases / antagonists & inhibitors*
  • Amyloid Precursor Protein Secretases / metabolism
  • Amyloid beta-Peptides / metabolism
  • Animals
  • CHO Cells
  • Cricetinae
  • Cricetulus
  • Enzyme Activation / drug effects
  • Ginsenosides / chemistry
  • Ginsenosides / pharmacology*
  • Humans
  • Membrane Microdomains / drug effects
  • Membrane Microdomains / metabolism*
  • Mice
  • Mice, Transgenic
  • Minor Histocompatibility Antigens
  • Neurons / drug effects
  • Neurons / metabolism
  • Neurons / pathology
  • Phosphatidylinositol 4,5-Diphosphate / metabolism
  • Phosphotransferases (Alcohol Group Acceptor) / metabolism*
  • Presenilin-1 / metabolism*
  • Protein Binding / drug effects
  • Protein Structure, Tertiary
  • Receptors, Nerve Growth Factor / chemistry
  • Receptors, Nerve Growth Factor / metabolism
  • Receptors, Notch / chemistry
  • Receptors, Notch / metabolism
  • Structure-Activity Relationship

Substances

  • Amyloid beta-Peptides
  • Ginsenosides
  • Minor Histocompatibility Antigens
  • Phosphatidylinositol 4,5-Diphosphate
  • Presenilin-1
  • Receptors, Nerve Growth Factor
  • Receptors, Notch
  • Ngfr protein, mouse
  • ginsenoside Rg3
  • Phosphotransferases (Alcohol Group Acceptor)
  • phosphatidylinositol phosphate 4-kinase
  • Amyloid Precursor Protein Secretases