The activation and differential signalling of the growth hormone receptor induced by pGH or anti-idiotypic monoclonal antibodies in primary rat hepatocytes

Mol Cell Endocrinol. 2013 Aug 25;376(1-2):51-9. doi: 10.1016/j.mce.2013.06.008. Epub 2013 Jun 14.

Abstract

In this report, we have developed a panel of monoclonal anti-idiotypic antibodies to pGH by immunising BALB/c mice with a purified monoclonal anti-pGH antibody (1A3), among which one mAb, termed CG-8F, was selected for further characterisation. We found that CG-8F behaved as a typical Ab2β, not only conformationally competing with pGH for 1A3 but also exhibiting recognition for GHR in a rat hepatocyte model. We next examined the resulting signal transduction pathways triggered by this antibody in rat hepatocytes and found that both pGH and CG-8F could trigger the JAK2-STAT1/3/5-mediated signal transduction pathway. Furthermore, the phosphorylation kinetics of pSTAT1/3/5 induced by either pGH or CG-8F were remarkably similar in the dose-response and time course rat hepatocyte experiments. In contrast, only pGH, but not CG-8F, was capable of inducing ERK phosphorylation. Further experimental studies indicated that the two functional binding sites on CG-8F are required for GHR activation. This study partially reveals the mechanism of action of GH anti-idiotypic antibodies and also indicates that monoclonal anti-idiotypic antibodies represent an effective way to produce GH mimics, suggesting that it is possible to produce signal-specific cytokine agonists using an anti-idiotypic antibody approach.

Keywords: Ab2; Anti-idiotypic monoclonal antibody; CLSM; FITC; GHR; Growth hormone; JAK; Janus kinase; MAPK; PBS; PRL; STAT; Signal transduction; TMB; TPO; anti-idiotypic antibody; confocal laser scanning microscope; fluorescein isothiocyanate; growth hormone receptor; mAb; mitogen-activated protein kinase; monoclonal antibody; pGH; phosphate-buffered saline; porcine growth hormone; prolactin; signal transducers and activators of transcription; tetramethylbenzidine; thrombopoietin.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Animals
  • Animals, Newborn
  • Antibodies, Anti-Idiotypic / biosynthesis
  • Antibodies, Anti-Idiotypic / isolation & purification
  • Antibodies, Anti-Idiotypic / pharmacology*
  • Antibodies, Monoclonal / biosynthesis
  • Antibodies, Monoclonal / isolation & purification
  • Antibodies, Monoclonal / pharmacology*
  • Gene Expression Regulation
  • Growth Hormone / metabolism*
  • Growth Hormone / pharmacology
  • Hepatocytes / cytology
  • Hepatocytes / drug effects*
  • Hepatocytes / metabolism
  • Janus Kinase 2 / genetics
  • Janus Kinase 2 / metabolism
  • Liver / cytology
  • Liver / drug effects*
  • Liver / metabolism
  • Mice
  • Mice, Inbred BALB C
  • Phosphorylation
  • Primary Cell Culture
  • Rats
  • Rats, Wistar
  • Receptors, Somatotropin / genetics*
  • Receptors, Somatotropin / metabolism
  • STAT1 Transcription Factor / genetics
  • STAT1 Transcription Factor / metabolism
  • STAT3 Transcription Factor / genetics
  • STAT3 Transcription Factor / metabolism
  • STAT5 Transcription Factor / genetics
  • STAT5 Transcription Factor / metabolism
  • Signal Transduction / drug effects*

Substances

  • Antibodies, Anti-Idiotypic
  • Antibodies, Monoclonal
  • Receptors, Somatotropin
  • STAT1 Transcription Factor
  • STAT3 Transcription Factor
  • STAT5 Transcription Factor
  • Stat1 protein, rat
  • Stat3 protein, rat
  • Growth Hormone
  • Jak2 protein, rat
  • Janus Kinase 2