Dissection of the IgE and T-cell recognition of the major group 5 grass pollen allergen Phl p 5

J Allergy Clin Immunol. 2014 Mar;133(3):836-45.e11. doi: 10.1016/j.jaci.2013.08.038. Epub 2013 Oct 31.

Abstract

Background: The major timothy grass pollen allergen Phl p 5 belongs to the most potent allergens involved in hay fever and asthma.

Objective: This study characterized immune-dominant IgE- and T-cell-recognition sites of Phl p 5.

Methods: Seven peptides, P1 to P7 with a length of 31 to 38 amino acids that spanned the Phl p 5 sequence, were synthesized, characterized by circular dichroism spectroscopy, and tested for IgE reactivity, basophil activation, and T-cell reactivity. Carrier-bound peptides were studied for their ability to induce IgG antibodies in rabbits which recognize Phl p 5 or cross-reactive allergens from different grass species. Peptide-specific antibodies were tested for the capability to inhibit IgE reactivity to Phl p 5 and allergen-induced basophil activation of patients with allergy.

Results: The peptides exhibited no secondary structure and showed no IgE reactivity or relevant allergenic activity, indicating that Phl p 5 IgE epitopes are conformational. Except for P3, peptide-specific IgG antibodies blocked IgE binding to Phl p 5 of patients with allergy and cross-reacted with temperate grasses. IgE inhibition experiments and molecular modeling identified several clustered conformational IgE epitopes on the N- as well as C-terminal domain of Phl p 5. P4, which stimulated the strongest T-cell and cytokine responses in patients, was not part of the major IgE-reactive regions.

Conclusion: Our study shows an interesting dissociation of the major IgE- and T-cell-reactive domains in Phl p 5 which provides a basis for the development of novel forms of immunotherapy that selectively target IgE or T-cell responses.

Keywords: Allergy; epitopes; grass pollen allergen; peptides.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Adolescent
  • Adult
  • Amino Acid Sequence
  • Animals
  • Basophils / physiology
  • Child
  • Cross Reactions
  • Cytokines / biosynthesis
  • Epitope Mapping
  • Female
  • Humans
  • Immunoglobulin E / immunology*
  • Lymphocyte Activation
  • Male
  • Middle Aged
  • Models, Molecular
  • Molecular Sequence Data
  • Plant Proteins / chemistry
  • Plant Proteins / immunology*
  • Protein Structure, Secondary
  • Rabbits
  • T-Lymphocytes / immunology*

Substances

  • Cytokines
  • Phl p V protein, Phleum pratense
  • Plant Proteins
  • Immunoglobulin E