1 Royal Belgian Institute of Natural Sciences, OD Taxonomy and Phylogeny (JEMU), Vautierstraat 29 - B-1000 Brussels, Belgium.
2 Royal Belgian Institute of Natural Sciences, OD Taxonomy and Phylogeny (JEMU), Vautierstraat 29 - B-1000 Brussels, Belgium ; Evolutionary Ecology Group, University of Antwerp, Groenenborgerlaan 171 - B-2020 Antwerp, Belgium.
3 Royal Museum for Central Africa (JEMU), Leuvensesteenweg 13 - B-3080 Tervuren, Belgium.
4 Evolutionary Ecology Group, University of Antwerp, Groenenborgerlaan 171 - B-2020 Antwerp, Belgium ; Royal Museum for Central Africa (JEMU), Leuvensesteenweg 13 - B-3080 Tervuren, Belgium.