The Kto-Skd complex can regulate ptc expression by interacting with Cubitus interruptus (Ci) in the Hedgehog signaling pathway

J Biol Chem. 2014 Aug 8;289(32):22333-41. doi: 10.1074/jbc.M114.560995. Epub 2014 Jun 24.

Abstract

The hedgehog (Hh) signaling pathway plays a very important role in metazoan development by controlling pattern formation. Drosophila imaginal discs are subdivided into anterior and posterior compartments that derive from adjacent cell populations. The anterior/posterior (A/P) boundaries, which are critical to maintaining the position of organizers, are established by a complex mechanism involving Hh signaling. Here, we uncover the regulation of ptc in the Hh signaling pathway by two subunits of mediator complex, Kto and Skd, which can also regulate boundary location. Collectively, we provide further evidence that Kto-Skd affects the A/P-axial development of the whole wing disc. Kto can interact with Cubitus interruptus (Ci), bind to the Ci-binding region on ptc promoter, which are both regulated by Hh signals to down-regulate ptc expression.

Keywords: Development; Drosophila; Hedgehog Signaling Pathway; Signal Transduction; Transcription.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Animals
  • Animals, Genetically Modified
  • Body Patterning
  • DNA-Binding Proteins / metabolism*
  • Drosophila Proteins / genetics*
  • Drosophila Proteins / metabolism*
  • Drosophila melanogaster / genetics
  • Drosophila melanogaster / growth & development*
  • Drosophila melanogaster / metabolism*
  • Eye Proteins / genetics
  • Eye Proteins / metabolism*
  • Gene Expression Regulation, Developmental
  • Genes, Insect
  • Hedgehog Proteins / genetics
  • Hedgehog Proteins / metabolism*
  • Multiprotein Complexes / metabolism
  • Mutation
  • Promoter Regions, Genetic
  • Receptors, Cell Surface / genetics*
  • Signal Transduction
  • Transcription Factors / metabolism*
  • Wings, Animal / growth & development
  • Wings, Animal / metabolism

Substances

  • DNA-Binding Proteins
  • Drosophila Proteins
  • Eye Proteins
  • Hedgehog Proteins
  • KTO protein, Drosophila
  • Multiprotein Complexes
  • Receptors, Cell Surface
  • Transcription Factors
  • ci protein, Drosophila
  • ptc protein, Drosophila
  • skd protein, Drosophila
  • hh protein, Drosophila