1 Department of Geography, University of Hawai'i at Mānoa, Honolulu, Hawaii 96822, USA.
2 Department of Biology, University of Hawaii at Mānoa, Honolulu, Hawaii 96822, USA.
3 1] Department of Biology, University of Hawaii at Mānoa, Honolulu, Hawaii 96822, USA [2] Hawaii Institute of Marine Biology, University of Hawaii at Mānoa, Kāne'ohe, Hawaii 96744, USA.
4 1] Hawaii Institute of Marine Biology, University of Hawaii at Mānoa, Kāne'ohe, Hawaii 96744, USA [2] Department of Oceanography, University of Hawaii at Mānoa, Honolulu, Hawaii 96822, USA.
5 1] Department of Geography, University of Hawai'i at Mānoa, Honolulu, Hawaii 96822, USA [2] Hawaii Institute of Marine Biology, University of Hawaii at Mānoa, Kāne'ohe, Hawaii 96744, USA.
6 1] Hawaii Institute of Marine Biology, University of Hawaii at Mānoa, Kāne'ohe, Hawaii 96744, USA [2] Trans-disciplinary Organization for Subtropical Island Studies (TRO-SIS), University of the Ryukyus, Senbaru, Nishihara, Okinawa 903-0213, Japan.