IGF-2R-Gαq signaling and cardiac hypertrophy in the low-birth-weight lamb

Am J Physiol Regul Integr Comp Physiol. 2015 Apr 1;308(7):R627-35. doi: 10.1152/ajpregu.00346.2014. Epub 2015 Jan 28.

Abstract

The cardiac insulin-like growth factor 2 receptor (IGF-2R) can induce cardiomyocyte hypertrophy in a heterotrimeric G protein receptor-coupled manner involving αq (Gαq) or αs (Gαs). We have previously shown increased left ventricular weight and cardiac IGF-2 and IGF-2R gene expression in low-birth-weight (LBW) compared with average-birth-weight (ABW) lambs. Here, we have investigated the cardiac expression of IGF-2 gene variants, the degree of histone acetylation, and the abundance of proteins in the IGF-2R downstream signaling pathway in ABW and LBW lambs. Samples from the left ventricle of ABW and LBW lambs were collected at 21 days of age. There was increased phospho-CaMKII protein with decreased HDAC 4 abundance in the LBW compared with ABW lambs. There was increased GATA 4 and decreased phospho-troponin I abundance in LBW compared with ABW lambs, which are markers of pathological cardiac hypertrophy and impaired or reduced contractility, respectively. There was increased histone acetylation of H3K9 at IGF-2R promoter and IGF-2R intron 2 differentially methylated region in the LBW lamb. In conclusion, histone acetylation of IGF-2R may lead to increased IGF-2R mRNA expression and subsequently mediate Gαq signaling early in life via CaMKII, resulting in an increased risk of left ventricular hypertrophy and cardiovascular disease in adult life.

Keywords: histone acetylation; insulin-like growth factor 2 receptor; low birth weight.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Acetylation
  • Age Factors
  • Animals
  • Animals, Newborn
  • Birth Weight*
  • Calcium-Calmodulin-Dependent Protein Kinase Type 2 / metabolism
  • GATA4 Transcription Factor / metabolism
  • GTP-Binding Protein alpha Subunits, Gq-G11 / metabolism*
  • Gene Expression Regulation
  • Heart Ventricles / metabolism*
  • Heart Ventricles / pathology
  • Heart Ventricles / physiopathology
  • Histone Deacetylases / metabolism
  • Histones / metabolism
  • Hypertrophy, Left Ventricular / genetics
  • Hypertrophy, Left Ventricular / metabolism*
  • Hypertrophy, Left Ventricular / pathology
  • Hypertrophy, Left Ventricular / physiopathology
  • Myocardial Contraction
  • Myocardium / metabolism*
  • Myocardium / pathology
  • Phosphorylation
  • Promoter Regions, Genetic
  • RNA, Messenger / metabolism
  • Receptor, IGF Type 2 / genetics
  • Receptor, IGF Type 2 / metabolism*
  • Sheep
  • Signal Transduction*
  • Troponin I / metabolism
  • Ventricular Function, Left

Substances

  • GATA4 Transcription Factor
  • Histones
  • RNA, Messenger
  • Receptor, IGF Type 2
  • Troponin I
  • Calcium-Calmodulin-Dependent Protein Kinase Type 2
  • Histone Deacetylases
  • GTP-Binding Protein alpha Subunits, Gq-G11