Endogenous Ligand for GPR120, Docosahexaenoic Acid, Exerts Benign Metabolic Effects on the Skeletal Muscles via AMP-activated Protein Kinase Pathway

J Biol Chem. 2015 Aug 14;290(33):20438-47. doi: 10.1074/jbc.M115.657379. Epub 2015 Jul 1.

Abstract

Docosahexaenoic acid (DHA) is an endogenous ligand of G protein-coupled receptor 120 (GPR120). However, the mechanisms underlying DHA action are poorly understood. In this study, DHA stimulated glucose uptake in the skeletal muscles in an AMP-activated protein kinase (AMPK)-dependent manner. GPR120-mediated increase in intracellular Ca(2+) was critical for DHA-mediated AMPK phosphorylation and glucose uptake. In addition, DHA stimulated GLUT4 translocation AMPK-dependently. Inhibition of AMPK and Ca(2+)/calmodulin-dependent protein kinase kinase blocked DHA-induced glucose uptake. DHA and GW9508, a GPR120 agonist, increased GPR120 expression. DHA-mediated glucose uptake was not observed in GPR120 knockdown conditions. DHA increased AMPK phosphorylation, glucose uptake, and intracellular Ca(2+) concentration in primary cultured myoblasts. Taken together, these results indicated that the beneficial metabolic role of DHA was attributed to its ability to regulate glucose via the GPR120-mediated AMPK pathway in the skeletal muscles.

Keywords: AMP-activated kinase (AMPK); diabetes; glucose transport; metabolism; signal transduction.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Adenylate Kinase / metabolism*
  • Animals
  • Cell Line
  • Docosahexaenoic Acids / metabolism*
  • Glucose / metabolism
  • Ligands
  • Mice
  • Muscle, Skeletal / enzymology
  • Muscle, Skeletal / metabolism*
  • Phosphorylation
  • Rats
  • Receptors, G-Protein-Coupled / metabolism*

Substances

  • FFAR4 protein, mouse
  • Ffar4 protein, rat
  • Ligands
  • Receptors, G-Protein-Coupled
  • Docosahexaenoic Acids
  • Adenylate Kinase
  • Glucose