Human RNase L tunes gene expression by selectively destabilizing the microRNA-regulated transcriptome

Proc Natl Acad Sci U S A. 2015 Dec 29;112(52):15916-21. doi: 10.1073/pnas.1513034112. Epub 2015 Dec 14.

Abstract

Double-stranded RNA (dsRNA) activates the innate immune system of mammalian cells and triggers intracellular RNA decay by the pseudokinase and endoribonuclease RNase L. RNase L protects from pathogens and regulates cell growth and differentiation by destabilizing largely unknown mammalian RNA targets. We developed an approach for transcriptome-wide profiling of RNase L activity in human cells and identified hundreds of direct RNA targets and nontargets. We show that this RNase L-dependent decay selectively affects transcripts regulated by microRNA (miR)-17/miR-29/miR-200 and other miRs that function as suppressors of mammalian cell adhesion and proliferation. RNase L mimics the effects of these miRs and acts as a suppressor of proliferation and adhesion in mammalian cells. Our data suggest that RNase L-dependent decay serves to establish an antiproliferative state via destabilization of the miR-regulated transcriptome.

Keywords: RNase L; adhesion; dsRNA; miR-200; microRNA.

Publication types

  • Research Support, N.I.H., Extramural
  • Research Support, Non-U.S. Gov't

MeSH terms

  • Animals
  • Blotting, Western
  • Cell Adhesion / genetics
  • Cell Line, Tumor
  • Cell Proliferation / genetics
  • Cells, Cultured
  • Endoribonucleases / genetics*
  • Endoribonucleases / metabolism
  • Gene Expression Regulation*
  • HeLa Cells
  • Humans
  • Mice, Knockout
  • MicroRNAs / genetics*
  • MicroRNAs / metabolism
  • Mutation
  • RNA Stability
  • RNA, Messenger / genetics
  • RNA, Messenger / metabolism
  • Reverse Transcriptase Polymerase Chain Reaction
  • Transcriptome*

Substances

  • MIRN17 microRNA, human
  • MIRN200 microRNA, human
  • MIRN29a microRNA, human
  • MicroRNAs
  • RNA, Messenger
  • Endoribonucleases
  • 2-5A-dependent ribonuclease