Inhibition of Acid Sphingomyelinase by Antidepressants Counteracts Stress-Induced Activation of P38-Kinase in Major Depression

Neurosignals. 2015;23(1):84-92. doi: 10.1159/000442606. Epub 2015 Dec 19.

Abstract

Background/aims: Major depressive disorder is a common disease with serious morbidity, including increased risk of death from suicide. Major depressive disorder is treated with antidepressants. However, the molecular targets of antidepressants remained ill-defined and require further elucidation.

Methods: Mice were treated with corticosterone to induce stress, amitriptyline and the p38-kinase (p38K) inhibitor SB239063 or a combination of these drugs. Phosphorylation of p38K in hippocampal neurons was determined by immunostaining with a phospho-specific antibody, neuronal proliferation using BrdU-labelling and behaviour employing a set of behavioural tests.

Results: Corticosterone induced phosphorylation/activation of p38K in the hippocampus in vivo. Antidepressants reversed the effect of corticosterone on p38K activation in wildtype mice, but had no effect in acid sphingomyelinase-deficient animals. Corticosterone also reduced neurogenesis and triggered depression-like behavioural changes, effects that were prevented by pharmacological inhibition of p38K.

Conclusion: Stress induces p38K phosphorylation/activation in the hippocampus and thereby reduces neurogenesis and induces depression-like symptoms, events that are prevented by antidepressants via inhibition of the acid sphingomyelinase/ceramide system.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Adaptation, Ocular / drug effects
  • Adaptation, Ocular / genetics
  • Amitriptyline / pharmacology
  • Amitriptyline / therapeutic use*
  • Animals
  • Antidepressive Agents / therapeutic use*
  • Corticosterone / toxicity
  • Disease Models, Animal
  • Enzyme Activation / drug effects
  • Enzyme Inhibitors / pharmacology
  • Exploratory Behavior / drug effects
  • Gene Expression Regulation, Enzymologic / drug effects
  • Gene Expression Regulation, Enzymologic / genetics
  • Hippocampus / drug effects
  • Imidazoles / pharmacology
  • Mice
  • Mice, Inbred C57BL
  • Mice, Transgenic
  • Neurogenesis / drug effects
  • Neurogenesis / genetics
  • Pyrimidines / pharmacology
  • Reaction Time / drug effects
  • Reaction Time / genetics
  • Sphingomyelin Phosphodiesterase / genetics
  • Sphingomyelin Phosphodiesterase / metabolism*
  • Sphingomyelins / pharmacokinetics
  • Stress, Psychological / chemically induced
  • Stress, Psychological / drug therapy*
  • Stress, Psychological / enzymology*
  • p38 Mitogen-Activated Protein Kinases / metabolism*

Substances

  • Antidepressive Agents
  • Enzyme Inhibitors
  • Imidazoles
  • Pyrimidines
  • Sphingomyelins
  • Amitriptyline
  • p38 Mitogen-Activated Protein Kinases
  • ASMase, mouse
  • Sphingomyelin Phosphodiesterase
  • SB 239063
  • Corticosterone