1 Department of Pathobiological Sciences, University of Wisconsin School of Veterinary Medicine, Madison, Wisconsin, USA.
2 Department of Biological Sciences, University of Maryland, Baltimore, Maryland, USA.
3 Department of Pathobiological Sciences, University of Wisconsin School of Veterinary Medicine, Madison, Wisconsin, USA Wisconsin National Primate Research Center, Madison, Wisconsin, USA.
4 Center for Clinical Epidemiology and Population Health, Marshfield Clinic Research Foundation, Marshfield, Wisconsin, USA.