Brown adipose tissue: Updates in cellular and molecular biology

Tissue Cell. 2016 Oct;48(5):452-60. doi: 10.1016/j.tice.2016.08.001. Epub 2016 Aug 8.

Abstract

Brown adipose tissue (BAT) is mainly composed of adipocytes, it is highly vascularized and innervated, and can be activated in adult humans. Brown adipocytes are responsible for performing non-shivering thermogenesis, which is exclusively mediated by uncoupling protein (UCP) -1 (a protein found in the inner mitochondrial membrane), the hallmark of BAT, responsible for the uncoupling of the proton leakage from the ATP production, therefore, generating heat (i.e. thermogenesis). Besides UCP1, other compounds are essential not only to thermogenesis, but also to the proliferation and differentiation of BAT, including peroxisome proliferator-activated receptor (PPAR) family, PPARgamma coactivator 1 (PGC1)-alpha, and PRD1-BF-1-RIZ1 homologous domain protein containing protein (PRDM) -16. The sympathetic nervous system centrally regulates thermogenesis through norepinephrine, which acts on the adrenergic receptors of BAT. This bound leads to the initialization of the many pathways that may activate thermogenesis in acute and/or chronic ways. In summary, this mini-review aims to demonstrate the latest advances in the knowledge of BAT.

Keywords: Brown adipocytes; Brown adipose tissue; Non-shivering thermogenesis; Obesity; Uncoupling protein 1.

Publication types

  • Review

MeSH terms

  • Adenosine Triphosphate / metabolism
  • Adipocytes / metabolism*
  • Adipose Tissue, Brown / growth & development*
  • Adipose Tissue, Brown / metabolism
  • Animals
  • Cell Differentiation / genetics*
  • Cell Proliferation / genetics
  • DNA-Binding Proteins / biosynthesis
  • DNA-Binding Proteins / genetics
  • Humans
  • Mice
  • Norepinephrine / metabolism
  • Peroxisome Proliferator-Activated Receptor Gamma Coactivator 1-alpha / biosynthesis
  • Peroxisome Proliferator-Activated Receptor Gamma Coactivator 1-alpha / genetics
  • Thermogenesis / genetics*
  • Transcription Factors / biosynthesis
  • Transcription Factors / genetics
  • Uncoupling Protein 1 / biosynthesis
  • Uncoupling Protein 1 / genetics

Substances

  • DNA-Binding Proteins
  • PRDM16 protein, human
  • Peroxisome Proliferator-Activated Receptor Gamma Coactivator 1-alpha
  • Ppargc1a protein, mouse
  • Transcription Factors
  • UCP1 protein, human
  • Uncoupling Protein 1
  • Adenosine Triphosphate
  • Norepinephrine