1 Program for Evolutionary Dynamics, Harvard University, Cambridge, MA; Centre for Applied Mathematics in Bioscience and Medicine, Montreal, QC, Canada.
2 Centre for Applied Mathematics in Bioscience and Medicine, Montreal, QC, Canada; Department of Mathematics and Statistics, McGill University, Montreal, QC, Canada; and.
3 Centre for Applied Mathematics in Bioscience and Medicine, Montreal, QC, Canada; Department of Physiology, McGill University, Montreal, QC, Canada.