1 Infectious Diseases Department, University Hospital Hotel-Dieu, Nantes University, Nantes, France.
2 Department of Pathology, University Hospital Hotel-Dieu, Nantes University, Nantes, France.
3 Faculté de Médecine, INSERM, Institut Cochin U1016, Laboratoire de Dermatologie-CNR Syphilis and AP-HP, Groupe Hospitalier Paris Centre Cochin-Hôtel Dieu-Broca, Service de Dermatologie-Vénéréologie, Université Sorbonne Paris Descartes, Paris, France.