1 Leeds Institute of Biomedical & Clinical Sciences, University of Leeds, St James's University Hospital, Leeds, UK.
2 Centre for Gastrointestinal Sciences, University of Manchester, Manchester, UK.
3 Gastroenterology and Hepatology, Mayo Clinic, Jacksonville, USA.
4 Division of Infection Immunity and Respiratory Medicine, School of Biological Sciences, University of Manchester, University Hospital of South Manchester, Manchester, UK.