[Nascent Polypeptide-Associated Complex as Tissue-Specific Cofactor during Germinal Cell Differentiation in Drosophila Testes]

Mol Biol (Mosk). 2017 Jul-Aug;51(4):677-682. doi: 10.7868/S0026898417040115.
[Article in Russian]

Abstract

During the process of spermatogenesis, the proliferation of spermatogonia (stem cell descendants) is replaced by their differentiation in growing spermatocytes responsible for the preparation to meiosis, which is accompanied by a cardinal change in transcriptional programs. We have demonstrated that, in drosophila, this process is accompanied by a splash of the expression of β-subunit of nascent polypeptide-associated complex (NAC) associated by ribosomes. Nascent polypeptide-associated complex is known as a chaperone involved in co-translational protein folding. This is the first case of the detection of tissue-specific co-translational NAC cofactor in multicellular eukaryotes. It is proposed that spermatocyte specific NAC is involved in the modulation of the expression of the proteins that provide the functioning of subsequent stages of spermatogenesis.

Keywords: spermatocytes; spermatogenesis; tissue-specific ribosomes; βNAC.

MeSH terms

  • Animals
  • Cell Differentiation
  • Drosophila Proteins / genetics*
  • Drosophila Proteins / metabolism
  • Drosophila melanogaster / genetics*
  • Drosophila melanogaster / growth & development
  • Drosophila melanogaster / metabolism
  • Gene Expression Regulation, Developmental*
  • Male
  • Meiosis
  • Molecular Chaperones / genetics*
  • Molecular Chaperones / metabolism
  • Protein Biosynthesis
  • Protein Subunits / genetics
  • Protein Subunits / metabolism
  • Ribosomes / genetics
  • Ribosomes / metabolism
  • Spermatocytes / growth & development
  • Spermatocytes / metabolism*
  • Spermatocytes / ultrastructure
  • Spermatogenesis / genetics
  • Testis / growth & development
  • Testis / metabolism*

Substances

  • Drosophila Proteins
  • Molecular Chaperones
  • Protein Subunits
  • nascent-polypeptide-associated complex